Cenchrus
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Cenchrus is a widespread genus of plants in the grass family, native to tropical and warm temperate regions of the world.[1][2] Its species are native to many countries in Asia, Africa, Australia, the Americas, and various oceanic islands.[3][4]
Common names include buffelgrasses, sandburs, and sand spur. Such names allude to the sharp, spine-covered burrs characterizing the inflorescences of the members of the genus.[5] Those previously classified as Pennisetum Template:IPAc-en[6] are known commonly as fountaingrasses (fountain grasses).[7][8][9][10]
Taxonomy
Pennisetum
Pennisetum is a former genus that heavily overlaps with Cenchrus,[11] and the boundary between them has been unclear.[12] Cenchrus was derived from Pennisetum and the two are grouped in a monophyletic clade.[13] A main morphological character that has been used to distinguish them is the degree of fusion of the bristles in the inflorescence, but this is often unreliable. In 2010, researchers proposed to transfer Pennisetum into Cenchrus, along with the related genus Odontelytrum.[14] The genus is currently not accepted as separate from Cenchrus in Kew's Plants of the World Online database.[15]
Distribution
The various species are native to Africa, Asia, Australia, and Latin America, with some of them widely naturalized in Europe and North America, as well as on various oceanic islands.[16]
Description
They are annual or perennial grasses. Some are petite while others can produce stems up to 8 meters tall.[12] The inflorescence is a very dense, narrow panicle containing fascicles of spikelets interspersed with bristles. There are three kinds of bristle, and some species have all three, while others do not. Some bristles are coated in hairs, sometimes long, showy, plumelike hairs that inspired the genus name, the Latin penna ("feather") and seta ("bristle").[12]
Uses
The genus includes pearl millet (P. glaucum), an important food crop. Napier grass (P. purpureum) is used for grazing livestock in Africa.
Several species are cultivated as ornamental plants, notably P. advena, P. alopecuroides, P. orientale, P. setaceum, and P. villosum. The cultivar 'Fairy Tails' is a recipient of the Royal Horticultural Society's Award of Garden Merit.[17][18]
Ecology
Many Pennisetum grasses are noxious weeds, including feathertop grass (P. villosum) and kikuyu grass (P. clandestinum), which is also a popular and hardy turf grass in some parts of the world.
The herbage and seeds of these grasses are food for herbivores, such as the chestnut-breasted mannikin (Lonchura castaneothorax), the caterpillar of the butterfly Melanitis phedima, and the larvae of the fly genus Delia.
The genus is a host of the pathogenic fungus Cochliobolus sativus.
Species
107 species are currently accepted.[15] They include:[19][20][21][22]
- Cenchrus abyssinicus (Hack.) Morrone – Ethiopia, Tanzania, Yemen, Limpopo, Mpumalanga
- Cenchrus agrimonioides Trin. – kāmanomano – Hawaiian Islands
- Cenchrus alopecuroides (L.) Thunb.Script error: No such module "Check for unknown parameters".
- Cenchrus americanus (L.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus annuus (Mez) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus arnhemicus (F.Muell.) Morrone – Australia (Western Australia and Northern Territory)
- Cenchrus articularis (Trin.) M.W.Tornab. & W.L.WagnerScript error: No such module "Check for unknown parameters".
- Cenchrus basedowii (Summerh. & C.E.Hubb.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus beckeroides (Leeke) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus biflorus Roxb. – kram-kram – Africa, Arabian Peninsula, Indian Subcontinent, and Madagascar
- Cenchrus brevisetosus (B.K.Simon) B.K.Simon – Australia (Western Australia, Northern Territory, and Queensland)
- Cenchrus brownii Roem. & Schult. – North America, South America, and West Indies
- Cenchrus cafer (Bory) VeldkampScript error: No such module "Check for unknown parameters".
- Cenchrus caliculatus Cav. – Australia, New Zealand, and assorted islands in the Pacific and Indian oceans
- Cenchrus caudatus (Schrad.) KuntzeScript error: No such module "Check for unknown parameters".
- Cenchrus chilensis (É.Desv.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus ciliaris L. – buffelgrass – Africa, Arabian Peninsula, Indian Subcontinent, and Sicily; naturalized as a nutritious arid pasture lands in parts of North and South America, Australia,[23] Southeast Asia, and various islands; considered noxious weed in some places
- Cenchrus clandestinus (Hochst. ex Chiov.) Morrone
- Cenchrus complanatus (Nees) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus × cupreus (Thorpe) GovaertsScript error: No such module "Check for unknown parameters".
- Cenchrus distachyus (E.Fourn.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus distichophyllus Griseb. – Cuba
- Cenchrus divisus (J.F.Gmel.) Verloove, Govaerts & ButtlerScript error: No such module "Check for unknown parameters".
- Cenchrus domingensis (Spreng. ex Schult.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus durus (Beal) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus echinatus L. – common sandbur – North America, South America, and West Indies; naturalized in parts of Africa, southern Asia, and various islands
- Cenchrus elegans (Hassk.) Veldkamp – Malesia
- Cenchrus elymoides F.Muell. – Australia (Western Australia, Northern Territory, and Queensland)
- Cenchrus exiguus (Mez) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus flaccidus (Griseb.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus flexilis (Mez) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus foermerianus (Leeke) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus geniculatus Thunb.Script error: No such module "Check for unknown parameters".
- Cenchrus glaucifolius (Hochst. ex A.Rich.) RudovScript error: No such module "Check for unknown parameters".
- Cenchrus gracilescens (Hochst.) ZonScript error: No such module "Check for unknown parameters".
- Cenchrus gracillimus Nash – West Indies and southeastern USA
- Cenchrus henryanus (F.Br.) M.W.Tornab. & W.L.WagnerScript error: No such module "Check for unknown parameters".
- Cenchrus hohenackeri (Hochst. ex Steud.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus hordeoides (Lam.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus intectus (Chase) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus lanatus (Klotzsch) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus latifolius (Spreng.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus laxius (Clayton) ZonScript error: No such module "Check for unknown parameters".
- Cenchrus ledermannii (Mez) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus longisetus M.C.Johnst.Script error: No such module "Check for unknown parameters".
- Cenchrus longispinus (Hack.) Fern. – mat sandbur – Canada, USA, and Mexico
- Cenchrus longissimus (S.L.Chen & Y.X.Jin) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus × longistylus (Hochst. ex A.Rich.) Thulin & S.M.PhillipsScript error: No such module "Check for unknown parameters".
- Cenchrus massaicus (Stapf) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus mezianus (Leeke) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus michoacanus H.F.Gut. & MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus mitis Andersson – Somalia, Kenya, Tanzania, Uganda, and Mozambique
- Cenchrus monostigma (Pilg.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus multiflorus J.Presl – Mexico, Central America
- Cenchrus mutilatus KuntzeScript error: No such module "Check for unknown parameters".
- Cenchrus myosuroides Kunth – North America, South America, and West Indies
- Cenchrus nanus (Engl.) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus nervosus (Nees) KuntzeScript error: No such module "Check for unknown parameters".
- Cenchrus nodiflorus (Franch.) ZonScript error: No such module "Check for unknown parameters".
- Cenchrus nubicus (Hochst.) ZonScript error: No such module "Check for unknown parameters".
- Cenchrus occidentalis (Chase) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus orientalis (Rich.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus palmeri Vasey – Baja California, Baja California Sur, Sonora, Sinaloa, and Arizona
- Cenchrus pennisetiformis Steud. – Africa, Arabian Peninsula, and Indian Subcontinent
- Cenchrus peruvianus (Trin.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus petiolaris (Hochst.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus pilcomayensis (Mez) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus pilosus Kunth – central Mexico to northern Chile
- Cenchrus pirottae (Chiov.) ZonScript error: No such module "Check for unknown parameters".
- Cenchrus platyacanthus Andersson – Galápagos
- Cenchrus prieurii (Kunth) Maire – Sahara, Arabian Peninsula, Indian Subcontinent, and Myanmar
- Cenchrus preslii (Kunth) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus procerus (Stapf) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus prolificus (Chase) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus pseudotriticoides (A.Camus) Voronts.Script error: No such module "Check for unknown parameters".
- Cenchrus pumilus (Hack.) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus purpureus (Schumach.) Morrone – Africa and Oman
- Cenchrus qianningensis (S.L.Zhong) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus ramosus (Hochst.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus rigidus (Griseb.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus riparius (Hochst. ex A.Rich.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus robustus R.D.Webster – Queensland and New South Wales
- Cenchrus rupestris (Chase) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus sagittatus (Henrard) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus schweinfurthii (Pilg.) ZonScript error: No such module "Check for unknown parameters".
- Cenchrus setaceus (Forssk.) MorroneScript error: No such module "Check for unknown parameters". – tender fountain grass, crimson fountaingrass – northern Africa and southwestern Asia; naturalized in Australia, New Zealand, and scattered places in Europe and the Americas
- Cenchrus setiger Vahl – Sahara, East Africa, Arabian Peninsula, Iran, Indian Subcontinent, Myanmar, and Andaman and Nicobar Islands
- Cenchrus shaanxiensis (S.L.Chen & Y.X.Jin) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus sichuanensis (S.L.Chen & Y.X.Jin) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus sieberianus (Schltdl.) VerlooveScript error: No such module "Check for unknown parameters".
- Cenchrus somalensis Clayton – Somalia
- Cenchrus sphacelatus (Nees) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus spinifex Cav. – coastal sandbur – North America, South America, and West Indies
- Cenchrus squamulatus (Fresen.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus stramineus (Peter) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus tempisquensis (R.W.Pohl) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus thulinii (S.M.Phillips) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus trachyphyllus (Pilg.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus tribuloides L. – sanddune sandbur – North America, South America, and West Indies
- Cenchrus trisetus (Leeke) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus tristachyus (Kunth) KuntzeScript error: No such module "Check for unknown parameters".
- Cenchrus uliginosus (Hack.) ined.Script error: No such module "Check for unknown parameters".
- Cenchrus unisetus (Nees) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus violaceus (Lam.) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus weberbaueri (Mez) MorroneScript error: No such module "Check for unknown parameters".
- Cenchrus yemensis (Deflers) Rudov & AkhaniScript error: No such module "Check for unknown parameters".
- Formerly included[19]
Several species are now considered better suited to other genera: Anthephora, Centotheca, Dactyloctenium, Echinaria, Echinolaena, Hackelochloa, Hilaria, Pennisetum, Phragmites, Scleria, Setaria, Trachys, Tragus, Tribolium.
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Sand spur in front of a centimeter scale
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Sandspurs with 1mm markings
See also
References
Further reading
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External links
Template:Sister project Template:Sister project
Template:Taxonbar Template:Authority control
- ↑ Cite error: Invalid
<ref>tag; no text was provided for refs nameds - ↑ Linnaeus, Carl von. 1753. Species Plantarum 2: 1049–1050 in Latin
- ↑ Flora of China Vol. 22 Page 552 蒺藜草属 ji li cao shu Cenchrus Linnaeus, Sp. Pl. 2: 1049. 1753.
- ↑ Altervista Flora Italiana, genere Cenchrus includes photos and range maps for several species
- ↑ Script error: No such module "citation/CS1".
- ↑ Sunset Western Garden Book. 1995. 606–07.
- ↑ Pennisetum. Integrated Taxonomic Information System (ITIS).
- ↑ Pennisetum. The Jepson eFlora 2013.
- ↑ Pennisetum. USDA PLANTS.
- ↑ Identified gaps for Pennisetum genepool. Template:Webarchive Crop Wild Relatives. CIAT.
- ↑ Martel, E., et al. (2004). Chromosome evolution of Pennisetum species (Poaceae): implications of ITS phylogeny. Plant Systematics and Evolution 249(3–4), 139–49.
- ↑ a b c Wipff, J. K. Pennisetum Rich. The Grass Manual. Flora of North America.
- ↑ Ozias-Akins, P., et al. (2003). Molecular characterization of the genomic region linked with apomixis in Pennisetum/Cenchrus. Functional & Integrative Genomics, 3(3), 94–104.
- ↑ Chemisquy, M. A., et al. (2010). Phylogenetic studies favour the unification of Pennisetum, Cenchrus and Odontelytrum (Poaceae): a combined nuclear, plastid and morphological analysis, and nomenclatural combinations in Cenchrus. Annals of Botany 106(1), 107–30.
- ↑ a b Cite error: Invalid
<ref>tag; no text was provided for refs namedpowo - ↑ Kew World Checklist of Selected Plant Families
- ↑ Script error: No such module "citation/CS1".
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- ↑ a b Script error: No such module "citation/CS1".
- ↑ Script error: No such module "template wrapper".
- ↑ Script error: No such module "citation/CS1".
- ↑ The Plant List search for Cenchrus
- ↑ Henry DR Pasture plants of Southern Inland Queensland. DPI. 1995