Bayanshiree Formation

From Wikipedia, the free encyclopedia
Jump to navigation Jump to search

Template:Short description Template:Lead too short Template:Infobox rockunit

The Bayanshiree Formation (also known as Baynshiree/Baynshire, Baynshirenskaya Svita, Baysheen Shireh, or Bayan Shireh[1]) is a geological formation in Mongolia, that dates to the Cretaceous period. It was first described and established by Vasiliev et al. 1959.[2]

Description

The Bayanshiree Formation is primarily composed by varicoloured claystones and sandstones with calcareous concretions and characterized by grey mudstones and yellowish-brown medium grained sandstones.[2][3] Up to Template:Convert thick, the most complete sections are found in the eastern Gobi Desert, consisting of fine-grained, often cross-stratified gray sandstone interbedded with claystone and concretionary, intraformational conglomerates with relatively thick units of red to brown mudstone in the upper part.[4][5] The Baynshire and Burkhant localities are mainly composed by mudstone, siltstone, sandstone, and conglomerates, with most of their sedimentation being fluvial.[6][7][8] The environments that were present on the Bayanshiree Formation consisted mainly on semi-arid climates with large water bodies, such as rivers or lakes.[9][8] Although it is considered to be partially lacustrine,[10] largescale cross-stratification in many of the sandstone layers at the Baynshire locality seem to indicate that a large meandering fluvial system was present. It has been implied that during the late Bayan Shirehnian times, large rivers with direct connections to the sea drained a prominent part of the eastern Gobi region.[4][7][9]

Stratigraphy

According to Jerzykiewicz and Russell, the Bayanshiree Formation can be divided into 2 informal units: upper and lower beds.[6] Danilov and colleagues have suggested that the lower beds are Cenomanian to early Turonian, and the upper beds are late Turonian to Santonian in age.[11] While the lower beds are composed by extensive conglomerate that indicates the ancient presence of very active rivers, the upper beds are mainly composed by mudstone and claystone that is interbedded by sandstone, indicating again, the presence of rivers but also lakes and other water bodies.[12]

Based on comparisons with other formations, Jerzykiewicz and Russell suggested that the Bayanshiree paleofauna seems to correspond best with the Turonian through early Campanian stages of the Late Cretaceous, about 93 to 80 million years ago.[6] However, examination of the magnetostratigraphy of the formation indicates that the entire Bayanshiree lies within the Cretaceous Long Normal, which lasted only until the end of the Santonian stage, giving a possible Cenomanian through Late Santonian age, between 98 and 83 million years ago.[7] In 2012, Averianov and Sues re-examined many formations from the Gobi Desert and using biostratigraphic occurrences and previous dating, the Bayanshiree Formation was considered to be Cenomanian to Santonian in age. The lower beds dating to 98 million and 90 million years ago (early Cenomanian to late Turonian), and the upper beds dating to 90 million and 83 million years ago (late Turonian to late Santonian).[9] Calcite U–Pb measurements performed by Kurumada and colleagues in 2020 have estimated the age of the Bayanshiree Formation between 95.9 ± 6.0 Ma and 89.6 ± 4.0 Ma.[13]

Correlations

A potential correlation between the Iren Dabasu Formation has been long suggested by most authors, mainly based on the highly similar vertebrate assemblages.[6][14][15][16] However, Van Itterbeeck et al. 2005 argued against this correlation concluding that instead, the Iren Dabasu Formation was coeval with the younger Nemegt Formation based on the charophyte and ostracode assemblages; therefore, these formations were dated to the Late Campanian-Early Maastrichtian.[17] Averianov and Sues instead proposed a correlation between the Bayanshiree, Iren Dabasu and Bissekty formations.[9] In 2015, Tsuihiji and team found the Iren Dabasu-Nemegt correlation to be inconsistent since the microfossil assemblages used by Van Itterbeeck and colleagues were not restricted to the Maastrichtian period and the similarities between these assemblages were most probably due to a similar deposition and climate settings.[18]

Paleobiota of the Bayanshiree Formation

In terms of biodiversity, therizinosaurs and turtles were the most abundant vertebrates across the formation, as evidenced on numerous remains.[11][19] Nevertheless, hadrosauroids were fairly abundant too, particularly at the Baynshire locality with numerous remains unearthed from this area and a new unnamed hadrosauroid. In addition, most specimens of Gobihadros come from this locality.[8][20][12] Also, niche partitioning has been reported within the therizinosaurids Erlikosaurus and Segnosaurus,[21] and the ankylosaurs Talarurus and Tsagantegia.[22] Mammals however, are extremely uncommon; Tsagandelta is the only mammal described so far.[23] Besides vertebrate fossils, abundant fossil fruits have been collected from the Bor Guvé and Khara Khutul localities and they are especially abundant at Bor Guvé. Although they resemble Abelmoschus esculentus their taxonomic position remains unclear and further examination is required.[24]

File:Bayan Shireh Formation Fauna.png
Bayanshiree Formation fauna

Template:Paleobiota-key-compact

Crocodylomorphs

Crocodylomorphs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images

Paralligator

P. gradilifrons Sheeregeen Gashoon "Complete skull with fragmentary postcrania."[25][26] A paralligatorid.
File:Paralligator major.jpg
P. major Sheeregeen Gashoon "Fragmentary crania."[27][26] A paralligatorid.
Unnamed crocodylomorph ichnotaxon Indeterminate Shine Us Khuduk "Swim tracks."[19] Tracks made underwater by a swimming individual.

Fish

Sharks reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Hybodus H. kansaiensis Bayshin Tsav "Teeth and postcrania."[9] A hybodontid.

File:Hybodus model.jpg

Osteichthyes spp. Indeterminate "Unspecified material."[9] Bony fish.

Flora

Flora reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Bothrocaryum B. gobience Khara Khutul "Unspecified material."[28] A cornacean.
Nyssoidea N. mongolica Khara Khutul "Unspecified material."[28] A cornacean.
Angiosperm spp. Indeterminate Bor Guvé, Khara Khutul "Fossil fruits."[24] Very abundant at Bor Guvé.
Taxodium T. sp[29] Khara Khutul A relative of the Bald Cypress

Mammals

Deltatheroidans reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Tsagandelta T. dashzevegi Tsagan Tsonj "Partial left dentary."[23] A deltatheroidan.
Bayshinoryctes B. shuvalovi "Dental remains"[30] A stem placental
Ravjaa R. ishiii "Lower jaw bones"[31] A zhelestid eutherian

Pterosaurs

Pterosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Azhdarchidae indet. Indeterminate Bayshin Tsav "Middle cervical vertebra."[32] An azhdarchid. File:Bayshin Tsav azhdarchid cervical vertebra.png
Azhdarchidae indet. Indeterminate Burkhant "Four cervical vertebrae."[32] An azhdarchid. File:Burkhant azhdarchid 3th cervical vertebra.png

Turtles

Turtles reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Charitonyx C. tajanikolaevae Khara Khutul "Fragmented postcrania."[33][34] A nanhsiungchelyid.
Gobiapalone G. orlovi Baynshire, Burkhant, Khongil Tsav "Skull and postcrania."[11] A trionychid.
Hanbogdemys H. orientalis Khara Khutul "Fragmented cervical remains."[34] A nanhsiungchelyid.
Kharakhutulia K. kalandadzei Khara Khutul Lower beds "Fragmented postcrania from multiple specimens."[34] A nanhsiungchelyid. File:Kharakhutulia holotype side views.png
Kirgizemys sp. Indeterminate Khara Khutul "Fragmented postcrania."[34] A nanhsiungchelyid.
Kizylkumemys K. schultzi Khara Khutul Lower beds "Fragmented postcrania."[35][34] A nanhsiungchelyid.
Lindholmemys L. martinsoni Sheeregeen Gashoon, Usheen Khuduk Upper beds "Fragmentary shell."[36] A lindholmemydid.
"Trionyx" T. baynshirensis Baynshire "Fragmentary shell."[11] A trionychine.
Pan-Carettochelys Indeterminate Uryl’b Usu Locality, Dornogovi Province Lower Beds Shell fragments A stem-carettochelyid, originally assigned to the species "Anosteira" shuwalovi[37]

Dinosaurs

Ornithischians

Ankylosaurs
Ankylosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Amtosaurus A. magnus Amtgai "Partial braincase." An ankylosaurid now regarded as nomen dubium.[38] File:Talarurus PIN 3780-1.jpg
Maleevus M. disparoserratus Sheeregeen Gashoon Upper beds "Partial maxillae and a referred braincase." An ankylosaurid now regarded as nomen dubium.[39]
Talarurus T. plicatospineus Bayshin Tsav, Baynshire, Shine Us Khuduk "Multiple specimens including partial to nearly complete skulls and postcranial remains."[40][41][42][43][22] An ankylosaurid. File:Talarurus.png
Tsagantegia T. longicranialis Tsagan Teg "Skull lacking lower jaws."[44][22] An ankylosaurid.
Ceratopsians
Ceratopsians reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Graciliceratops G. mongoliensis Sheeregeen Gashoon Upper beds "Fragmented skull and postcrania."[45][46] A neoceratopsian originally identified as Microceratus. File:Graciliceratops BW.jpg
Unnamed neoceratopsian Indeterminate Bayshin Tsav "Left maxilla and teeth."[47] A neoceratopsian.
Hadrosaurs
Hadrosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Amblydactylus spp. Indeterminate Baynshire "Five three-toed footprints."[48] Ornithopod tracks.
Gobihadros G. mongoliensis Bayshin Tsav, Baynshire, Char Teeg, Khongil Tsav, Khoorai Tsav Upper beds "Multiple specimens with a virtually complete skull and postcranial remains".[20][12] A hadrosauroid. File:Gobihadros skeleton.png
Hadrosauroidea indet. Indeterminate Shine Us Khuduk "Isolated surangular".[49] A hadrosauroid.
Ovaloolithidae indet. Indeterminate Mogoyn Ulagiyn Khaets "Eggs, shells and egg clutches."[50] Eggs probably laid by a hadrosaur.
Spheroolithus sp. Indeterminate Sheeregeen Gashoon "Egg clutch."[51] Eggs probably laid by a hadrosaur.
Undescribed Hadrosauroidea Indeterminate Amtgai, Bayshin Tsav, Bayshin Tsav IV, Khoorai Tsav "Multiple postcranial remains including an articulated skeleton."[8] Hadrosauroid remains.
Undescribed hadrosauroids Indeterminate Baynshire "Remains of at least two juvenile hadrosauroids."[52] Juvenile hadrosauroids in a caenagnathoid nesting site.
Unnamed hadrosauroid Indeterminate Baynshire, Char Teeg Upper beds "Partial right dentary and sparse postcranial remains."[12] A hadrosauroid slightly smaller than Gobihadros.
Unnamed hadrosauroid Indeterminate Bayshin Tsav "Nearly complete skeleton from a juvenile individual."[53] A hadrosauroid distinct from Gobihadros. File:Juvenile hadrosaur.jpg
Pachycephalosaurs
Pachycephalosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Amtocephale A. gobiensis Amtgai "Nearly complete frontoparietal dome."[54] A primitive pachycephalosaurid. File:Amtocephale LM.png

Sauropods

Sauropods reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Erketu E. ellisoni Bor Guvé "Vertebrae and postcrania."[24] A somphospondylan. File:Erketu cervical vertebrae.png
Sauropoda indet. Indeterminate Amtgai "Partial vertebrae and pelvic elements."[49] A sauropod.
Unnamed sauropod Indeterminate Shine Us Khuduk "Articulated skeleton."[49] A sauropod.
Unnamed sauropod ichnogenus Indeterminate Khavirgiin Dzo "Four consecutive hind footprints."[55] Sauropod tracks.
Unnamed titanosaur Indeterminate Bor Guvé "Vertebrae."[56] A titanosaur.

Theropods

Dromaeosaurids
Dromaeosaurids reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Achillobator A. giganticus Burkhant "Maxilla and fragmentary postcrania."[57] A giant dromaeosaurid. File:Achillobator reconstruction.png
Dromaeosauridae indet. Indeterminate Burkhant "Claw and large postcranial remains."[58][59] A dromaeosaurid.
Unnamed velociraptorine Indeterminate Shine Us Khuduk, Tel Ulan Chaltsai "Fragmentary crania and postcrania from two specimens."[60][61][62] A dromaeosaurid.
Ornithomimosaurs
Ornithomimosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
"Gallimimus" "G. mongoliensis" Bayshin Tsav "Complete skull with postcrania only lacking the caudal region."[63] An informally named ornithomimid that actually differs from Gallimimus. File:Gallimimus mongoliensis mount.jpg
Garudimimus G. brevipes Bayshin Tsav "Skull with fragmented postcrania."[64][65][66] An ornithomimosaur. File:Garudimimus Restoration.png
Undescribed ornithomimosaur Indeterminate Amtgai "Nearly Template:Convert long ulna."[67] A large ornithomimosaur.
Undescribed ornithomimosaurs Indeterminate Baynshire, Bayshin Tsav, Khongil Tsav, Sheeregeen Gashoon, Shine Us Khuduk "More than five specimens and a bonebed containing mainly postcranial elements."[49][68][69][70][71] Multiple ornithomimosaurian taxa.
Oviraptorosaurs
Oviraptorosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Caenagnathoidea indet. Indeterminate Baynshire "Nests including elongatoolithid eggs and associated juveniles."[52] A large caenagnathoid nesting site.
Macroelongatoolithus indet. Indeterminate Baynshire "Eggs within a large nest."[72] Eggs probably laid by a large oviraptorosaur.
Unnamed caenagnathid Indeterminate Tsagan Teg "Partial lower jaw similar to Gigantoraptor."[18] A large caenagnathid. File:LH V0011 & MPC-D 107 17 lower jaws.png
Therizinosaurs
Therizinosaurs reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Dendroolithidae spp. Indeterminate Baynshire "Various nests and egg clutches."[73][74] Eggs probably laid by a therizinosaur File:Segnosaurus nest 2.jpg
Duonychus[75] D. tsogtbaatari Urlibe Khudak Partial post-cranial skeleton A therizinosaurid

Duonychus_Restoration

Enigmosaurus E. mongoliensis Khara Khutul "Incompletely preserved pelvis."[76][77][78] A therizinosauroid. File:Enigmosaurus Restoration.jpg
Erlikosaurus E. andrewsi Bayshin Tsav Upper beds "Skull, right pes, and left humerus."[77][79][78] A therizinosaurid File:Erlikosaurus Restoration.png
Segnosaurus S. galbinensis Amtgai, Bayshin Tsav, Khara Khutul, Urlibe Khudak "Multiple specimens including the lower jaws and partial postcrania."[80][77][78] A large therizinosaurid File:Segnosaurus Restoration.jpg
Undescribed therizinosaurs Indeterminate Bayshin Tsav, Khara Khutul, Shine Us Khuduk "Cranial and postcranial elements from multiple specimens."[81][82][58][83][84][49][78] Some elements may represent already named taxa.
Troodontids
Troodontids reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Undescribed troodontid Indeterminate Bayshin Tsav "Partial vertebrae, forelimbs and hindlimbs."[85][86] A troodontid.
Tyrannosauroids
Tyrannosauroids reported from the Bayanshiree Formation
Genus Species Location Stratigraphic Position Material Notes Images
Khankhuuluu[87] K. mongoliensis Three specimens, consisting of partial cranial and postcranial material Bayshin Tsav specimens originally identified as Alectrosaurus File:Khankhuuluu mongoliensis.png
Other Theropods
Genus Species Location Stratigraphic Position Material Notes Images
Deinonychosauria indet. Indeterminate Shine Us Khuduk "Isolated remains including a pedal phalanx II-2."[49] The pedal phalanx II-2 is similar to Achillobator.
Theropoda indet. Indeterminate Urlibe Khudak "Nest with eggs."[8] Eggs probably laid by a theropod.

See also

References

Template:Reflist

  1. Cite error: Invalid <ref> tag; no text was provided for refs named benton2000
  2. a b Script error: No such module "citation/CS1".
  3. Script error: No such module "Citation/CS1".
  4. a b Script error: No such module "Citation/CS1".
  5. Script error: No such module "Citation/CS1".
  6. a b c d Script error: No such module "Citation/CS1".
  7. a b c Script error: No such module "Citation/CS1".
  8. a b c d e Script error: No such module "Citation/CS1".
  9. a b c d e f Script error: No such module "Citation/CS1".
  10. Script error: No such module "Citation/CS1".
  11. a b c d Script error: No such module "Citation/CS1".
  12. a b c d Script error: No such module "Citation/CS1".
  13. Script error: No such module "Citation/CS1".
  14. Script error: No such module "Citation/CS1".
  15. Script error: No such module "citation/CS1". Translated paper
  16. Script error: No such module "Citation/CS1".
  17. Script error: No such module "Citation/CS1".
  18. a b Script error: No such module "Citation/CS1".
  19. a b Script error: No such module "Citation/CS1".
  20. a b Script error: No such module "Citation/CS1".
  21. Script error: No such module "Citation/CS1".
  22. a b c Script error: No such module "Citation/CS1".
  23. a b Script error: No such module "Citation/CS1".
  24. a b c Script error: No such module "Citation/CS1".
  25. Script error: No such module "Citation/CS1".
  26. a b Script error: No such module "Citation/CS1".
  27. Script error: No such module "Citation/CS1".
  28. a b Script error: No such module "citation/CS1".
  29. Script error: No such module "Citation/CS1".
  30. Script error: No such module "Citation/CS1".
  31. Script error: No such module "Citation/CS1".
  32. a b Script error: No such module "Citation/CS1".
  33. Script error: No such module "Citation/CS1".
  34. a b c d e Script error: No such module "Citation/CS1".
  35. Script error: No such module "citation/CS1".
  36. Script error: No such module "Citation/CS1".
  37. Script error: No such module "Citation/CS1".
  38. Script error: No such module "Citation/CS1".
  39. Template:Cite thesis
  40. Script error: No such module "Citation/CS1".
  41. Script error: No such module "Citation/CS1".
  42. Script error: No such module "citation/CS1".
  43. Script error: No such module "Citation/CS1".
  44. Script error: No such module "Citation/CS1".
  45. Script error: No such module "Citation/CS1".
  46. Script error: No such module "citation/CS1".
  47. Script error: No such module "Citation/CS1".
  48. Script error: No such module "Citation/CS1".
  49. a b c d e f Script error: No such module "Citation/CS1".
  50. Script error: No such module "citation/CS1".
  51. Script error: No such module "Citation/CS1".
  52. a b Script error: No such module "Citation/CS1".
  53. Script error: No such module "Citation/CS1".
  54. Script error: No such module "Citation/CS1".
  55. Script error: No such module "Citation/CS1".
  56. Script error: No such module "Citation/CS1".
  57. Script error: No such module "Citation/CS1".
  58. a b Script error: No such module "Citation/CS1".
  59. Script error: No such module "Citation/CS1".
  60. Script error: No such module "Citation/CS1".
  61. Script error: No such module "Citation/CS1".
  62. Template:Cite thesis
  63. Script error: No such module "Citation/CS1".
  64. Script error: No such module "Citation/CS1".
  65. Script error: No such module "Citation/CS1".
  66. Script error: No such module "Citation/CS1".
  67. Script error: No such module "Citation/CS1".
  68. Script error: No such module "Citation/CS1".
  69. Script error: No such module "Citation/CS1".
  70. Script error: No such module "Citation/CS1".
  71. Template:Cite thesis
  72. Script error: No such module "Citation/CS1".
  73. Script error: No such module "Citation/CS1".
  74. Script error: No such module "Citation/CS1".
  75. Script error: No such module "Citation/CS1".
  76. Script error: No such module "Citation/CS1".
  77. a b c Script error: No such module "Citation/CS1".
  78. a b c d Script error: No such module "Citation/CS1".
  79. Script error: No such module "Citation/CS1".
  80. Script error: No such module "Citation/CS1".
  81. Script error: No such module "Citation/CS1".
  82. Script error: No such module "Citation/CS1".
  83. Script error: No such module "Citation/CS1".
  84. Script error: No such module "Citation/CS1".
  85. Script error: No such module "Citation/CS1".
  86. Script error: No such module "Citation/CS1".
  87. Script error: No such module "Citation/CS1".
  88. Script error: No such module "Citation/CS1".
  89. Script error: No such module "Citation/CS1".
  90. Script error: No such module "Citation/CS1".