Staffordiidae

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Staffordiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea.[1]

Staffordiidae is the only family in the superfamily Staffordioidea. This family has no subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Staffordiidae is a poorly understood[2] family, because it occurs only in the Dafla Hills area of India. The fauna and flora of that area has not been researched sufficiently.[2]

Various sources consider the family Staffordiidae as part of Dyakiidae[3] or Ariophantidae/Dyakiinae.[4]

Distribution

The distribution of the Staffordiidae includes only India in the Dafla Hills.[2]

This area is close to northern margin of the Indian Plate.[2] The historical area of origin of the Staffordiidae has not been researched because the coastal area in southern Asia where it is found became uninhabitable[2] after the Indian Plate and Eurasian Plate collided 50 to 55 million years ago. The original ancestral area of limacoid families is thought to be the Palaearctic region and south-eastern Asia.[2] Thus, it has been hypothesized that the Staffordiidae colonized its current area from the southern margin of the Asian part of the Eurasian Plate during the Oligocene period.[2]

Genera

Genera within the family Staffordiidae include:

The generic name Staffordia is in honor of Brigadier-General Stafford,Script error: No such module "Unsubst". who was in command of the punitive force which entered the Dafla Hills for the first time in the winter of 1874–1875.[5]

The foot of Staffordia is pointed.[5] The peripodial margin is simple with a narrow pale margin.[5] There are small right and left shell-lobes.[5]

Reproductive system of Staffordia: the dart-sac is small, globose, with a long cord-like attachment to a coronal gland.[5] The penis is simple.[5] The spermatheca is long.[5]

The radula of Staffordia has aculeate lateral teeth.[5]

Comparison of shells of three Staffordia species:

File:Staffordia daflaensis shell.png
Staffordia daflaensis
File:Staffordia staffordi shell.png
type specimen of Staffordia staffordi is juvenile
File:Staffordia toruputuensis shell.png
type specimen of Staffordia toruputuensis is juvenile

Cladogram

Staffordiidae is considered a sister group of all other families in the limacoid clade.[2]

The following cladogram shows the phylogenic relationships of this family and superfamily to the other families within the limacoid clade:[2]

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References

This article incorporates public domain text from the reference.[5]

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  1. MolluscaBase eds. (2021). MolluscaBase. Staffordiidae Thiele, 1931. Accessed through: World Register of Marine Species at: http://marinespecies.org/aphia.php?p=taxdetails&id=994727 on 2021-02-22
  2. a b c d e f g h i Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379–390. Script error: No such module "CS1 identifiers"., JSTOR.
  3. Barker G. M. (2001) Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. 1–146. In: Barker G. M. (ed.) (2001) The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, cited pages: 139–144. Template:ISBN.
  4. a b c d Ramakrishna, Dey A. & Mitra S. C. (PDF created 6 April 2010). "Checklist of Indian Land Mollusca" Template:Webarchive. Zoological Survey of India. accessed 30 June 2010. 65 pp.
  5. a b c d e f g h i j k Godwin-Austen H. H. (1907). Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean; Supplementary to Masers Theobald and Hanley's Conchologica Indica. Taylor and Francis, London. 2: page 184, plate CXIII.

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  • Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

Further reading

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  • Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309–1466.

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