Istiodactylidae

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Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus Istiodactylus was discovered not to be a member of the genus Ornithodesmus.

Systematics and distribution

File:Istiodactylid skulls.tif
Istiodactylid skulls: A) Istiodactylus latidens (NHMUK R3877) (B) Istiodactylus sinensis (NGMC 99-07-11) and (C) Nurhachius ignaciobritoi (IVPP V-13288)

Remains of taxa that can be confidently assigned to Istiodactylidae have been found in the UK and China, in rocks dating from the Early Cretaceous period (Barremian to Aptian stage).[1] Arbour and Currie (2011) described Canadian Gwawinapterus beardi as a member of Istiodactylidae living in the late Cretaceous (upper Campanian stage);[2] however, Witton (2012) suggested the tooth replacement pattern in this animal does not match that of pterosaurs, suggesting that the species might be non-pterosaurian.[1] Additional research suggested that the species was in fact a fish.[3] The earliest known species might be Archaeoistiodactylus linglongtaensis, from the Middle Jurassic of China;[4] however, it also has been suggested that the holotype specimen of this species might actually be a poorly preserved specimen of Darwinopterus.[5] Hongshanopterus, a supposed istiodactylid from China, has been reclassified as a non-istiodactylid member of Ornithocheiroidea of uncertain phylogenetic placement by Witton (2012).[1]

Istiodactylids were medium-sized pterosaurs with flat, rounded jaws similar to that of a duck. They had small teeth lining their jaws, however, and this can mostly be seen in the more advanced genera such as Istiodactylus.[6]

Classification

File:IstiodactylusDB3.jpg
Restoration of an Istiodactylus in flight

Below is a cladogram showing the phylogenetic placement of this group within Pteranodontia from Andres and Myers (2013).[7]

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The cladogram below is a topology recovered by Kellner et al. (2019). In the analyses, they recovered Istiodactylidae within the more inclusive group Istiodactyliformes, and assigned both Istiodactylus and Liaoxipterus to a new subfamily called Istiodactylinae, but kept Nurhachius as a basal member.[8] Script error: No such module "Clade".

Paleobiology

Lifestyle

Unlike most ornithocheiroids, istiodactylids bear morphologies suited to a terrestrial life and many of their fossils have been found in freshwater-deposits. Istiodactylids are considered to be pterosaurian equivalents to vultures: acting as the clean-up crew in their habitats. The swimming abilities of Istiodactylids have not been analyzed, in contrast to Ornithocheirids.Script error: No such module "Unsubst".

References

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