Coulter pine

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Coulter pine (Pinus coulteri), or big-cone pine, is a conifer in the genus Pinus of the family Pinaceae. Coulter pine is an evergreen conifer that lives up to 100 years. [1] It is a native of the coastal mountains of Southern California in the United States and northern Baja California in Mexico, occurring in mediterranean climates, where winter rains are infrequent and summers are dry with occasional thunderstorms.[2] Isolated groves are found as far north as Clearlake, California, on the flanks of Mt. Konocti and in Black Diamond Mines Regional Preserve. Although geographically isolated, these Coulter pine populations were very similar in all of three studies of morphological characteristics. Oleoresins (volatile portions) were also similar.[3]

While the species has a limited range in the wild, the Coulter pine is a popular ornamental tree and is grown in many countries.

Coulter pine (Pinus coulteri) is named after Thomas Coulter, an Irish botanist and physician. The Coulter pine produces some of the heaviest cones of any pine tree, up to Template:Cvt; among conifers, these are exceeded only by the cones of Araucaria bidwillii. Script error: No such module "Unsubst".

Description

Pinus coulteri is a substantial coniferous evergreen tree in the genus Pinus. The size ranges from Template:Cvt tall,Template:Sfn with a trunk diameter up to Template:Cvt. The trunk is vertical and branches horizontal to upcurved. The bark is dark gray-brown to near black, deeply furrowed, with long, scaly, irregularly anastomosing, rounded ridges.[4] The crown is pyramidal and may be dense or open, depending upon the site.[1] The leaves are needle-like, in bundles of three, glaucous gray-green in color, Template:Cvt long, and stout, Template:Cvt thick.Script error: No such module "Unsubst".

The exemplary characteristic of this tree is the large, spiny cones, which are Template:Cvt long (occasionally as much as Template:Cvt length has been observed)[5] and Template:Cvt in width,[6] and weigh Template:Cvt when fresh. Each segment, or "scale", of the cone is tipped with a talon-like hook. Although the slender cones of the sugar pine are longer, Coulter pines produce the largest cones of any pine species, and people are advised to wear hardhats when working in Coulter pine groves. The large size of the cones, combined with the claw-like scales, has earned them the nickname "widowmakers" among locals.Script error: No such module "Unsubst". Seed cones mature in two years, gradually shedding seeds thereafter, and are moderately persistent, massive, heavy, drooping, asymmetric at the base, narrowly ovoid before opening, ovoid-cylindric when open, Template:Cvt long, pale yellow-brown in color, and resinous, with stalks to Template:Cvt. The apophyses are transverse-rhombic, strongly and sharply cross-keeled, elongate, curved, and continuous with umbos to form long, upcurved claws Template:Cvt. The seeds are obovoid in shape, dark brown in color, and the body Template:Cvt long, with a wing to Template:Cvt.[4]

Taxonomy

The name Pinus coulteri comes from Latin for pine, and coulteri comes from its discoverer Thomas Coulter (1793–1843), an Irish botanist and physician.[7] Pinus coulteri was discovered by Coulter on the mountains of Santa Lucia, near the Mission of San Antonio, at latitude 36°, within sight of the ocean and at an elevation between 3000 and 4000 feet above sea level. It was growing intermingled with Pinus Lambertiana.[8]

This is a member of subsection Ponderosae, and is generally recognized as closely related (morphologically, geographically, and genetically) to Pinus sabiniana, P.Template:Nbsptorreyana, and P. jeffreyi; the three taxa form group Sabinianae Loudon 1838 (Pinus jeffreyi, though morphologically a bit different, is known to hybridize with P.Template:Nbspcoulteri, and molecular analysis clearly places it in the Sabinianae).[9] P. coulteri, alone in the Sabinianae, also shares a chloroplast haplotype with P. arizonica of the northern Sierra Madre. It seems likely that the species arose through a hybridization event involving two members of subsection Ponderosae.[9]

Distribution and habitats

Coulter pine is most frequent on steep south-facing slopes and ridges.[10][11][12][13] Soils may be poor to fertile, and are typically dry. Coulter pine is an indicator of serpentine soils, but also occurs on a variety of other substrates.  Soils range from loamy to gravelly or rocky in texture.[13][14][15] Coulter pine occurs at Template:Cvt elevation.[16][17]

Coulter pine is the dominant species in the following published classifications: Terrestrial natural communities of California,[13] Vegetation types of the San Bernardino Mountains,[18] Vegetation of the San Bernardino Mountains,[2] A vegetation classification system applied to southern California,[19] Mixed evergreen forest,[20] Vascular plant communities of California,[16] Montane and subalpine forests of the Transverse and Peninsular ranges.[21]

Ecology

The Coulter pine is closely related to the foothill pine, Pinus sabiniana. It is more distantly related to Jeffrey pine, with which it shares habitats, and the ponderosa pine.Template:Sfn Coulter pines tend to grow in drier environments than ponderosa and Jeffery pines.Script error: No such module "Unsubst".

This erect, medium-sized pine prefers south-facing slopes from Template:Cvt elevation, and tolerates dry rocky soil. Pinus coulteri most often appears in mixed forests. The Coulter pine occurs in a number of forest plant associations; for example, at higher elevations forestation of the San Jacinto Mountains Coulter Pine is co-dominant with the California black oak.Template:Sfn Woodpeckers often forage on the species, and peel the bark to access insects underneath.Template:Sfn

Uses

Wildlife, especially squirrels, gather the large seeds. The seeds were also once eaten by Native Americans.[22]

The wood is weak and soft, so that the species is little used other than for firewood.Script error: No such module "Unsubst".

Pinus coulteri is cultivated as an ornamental tree, planted in parks and large gardens, and used in drought-tolerant landscaping. The Coulter pine has gained the Royal Horticultural Society's Award of Garden Merit.Template:Sfn

Gallery

References

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Bibliography

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External links

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  1. ↑ a b Horton, Jerome S. 1949. Trees and shrubs for erosion control of southern California mountains. Berkeley, CA: U.S. Department of Agriculture, Forest Service, California [Pacific Southwest] Forest and Range Experiment Station; California Department of Natural Resources, Division of Forestry. 72 p. [10689]
  2. ↑ a b Minnich, Richard A. 1976. Vegetation of the San Bernardino Mountains. In: Latting, June, ed. Symposium proceedings: plant communities of southern California; 1974 May 4; Fullerton, CA. Special Publication No.2. Berkeley, CA: California Native Plant Society: 99-124.  [4232]
  3. ↑ St. John, Harold. 1973. List and summary of the flowering plants in the Hawaiian islands. Hong Kong: Cathay Press Limited. 519 p. [25354]
  4. ↑ a b Kral, R. 1993. Pinus. Flora of North America Editorial Committee (eds.): Flora of North America North of Mexico, Vol. 2. Oxford University Press.
  5. ↑ Script error: No such module "citation/CS1".
  6. ↑ Gardener's Chronicle Volume 23 (third series)(March 28, 1885) Page 415
  7. ↑ Trees of Stanford and Environs, by Ronald Bracewell, published 2005.
  8. ↑ Script error: No such module "Citation/CS1".
  9. ↑ a b Script error: No such module "Citation/CS1".
  10. ↑ Script error: No such module "citation/CS1".
  11. ↑ Barbour, Michael G. 1988. Californian upland forests and woodlands. In: Barbour, Michael G.; Billings, William Dwight, eds. North American terrestrial vegetation. Cambridge; New York: Cambridge University Press:131-164. [13880]
  12. ↑ Vogl, Richard J. 1976. An introduction to the plant communities of the Santa Ana and San Jacinto Mountains. In: Latting, June, ed. Symposium proceedings: plant communities of southern California; 1974 May 4; Fullerton, CA. Special Publication No. 2. Berkeley, CA: California Native Plant Society: 77-98. [4230]
  13. ↑ a b c Holland, Robert F. 1986. Preliminary descriptions of the terrestrial natural communities of California. Sacramento, CA: California Department of Fish and Game. 156 p. [12756]
  14. ↑ Krochmal, Arnold; Krochmal, Connie. 1982. Uncultivated nuts of the United States. Agriculture Information Bulletin 450. Washington, DC: U.S. Department of Agriculture, Forest Service. 89 p.  [1377]
  15. ↑ Script error: No such module "citation/CS1".
  16. ↑ a b Thorne, Robert F. 1976. The vascular plant communities of California. In: Latting, June, ed. Symposium proceedings: plant communities of southern California; 1974 May 4; Fullerton, CA. Special Publication No. 2. Berkeley, CA: California Native Plant Society: 1-31. [3289]
  17. ↑ Zobel, Bruce. 1953. Geographic range and intraspecific variation of Coulter pine. Madrono. 11(8): 285-316. [21797]
  18. ↑ Script error: No such module "citation/CS1".
  19. ↑ Template:Cite report
  20. ↑ Sawyer, John O.; Thornburgh, Dale A.; Griffin, James R. 1977. Mixed evergreen forest. In: Barbour, Michael G.; Major, Jack, eds. Terrestrial vegetation of California. New York: John Wiley and Sons: 359-381. [7218]
  21. ↑ Thorne, Robert F. 1977. Montane and subalpine forests of the Transverse and Peninsular ranges. In: Barbour, Michael G.; Major, Jack, eds. Terrestrial vegetation of California. New York: John Wiley and Sons: 537-557.  [7214]
  22. ↑ Script error: No such module "citation/CS1".
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