Chorrillo Formation

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File:Chorrillo Formation Dinosauria Size Comparison.svg
Non-avian dinosaurs named from the Chorrillo Formation

The Chorrillo Formation, also named as Chorillo Formation,[1] is a Maastrichtian (Late Cretaceous Epoch, 72.1–66 million years ago) geologic formation in southern Patagonia, Argentina. The formation is more than Script error: No such module "convert". thick and underlies the Calafate Formation and rests on top of the La Irene Formation.[2]

Fossils

Dinosaur remains are among the fossils that have been recovered from the formation.

Fossils recovered from the formation are:

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Dinosaurs

Ornithopods

Ornithopods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Euiguanodontia Indet.Script error: No such module "Unsubst". IndeterminateScript error: No such module "Unsubst". Proximal end of a femur Script error: No such module "Unsubst".
Hadrosauridae Indet.Script error: No such module "Unsubst". IndeterminateScript error: No such module "Unsubst". Two incomplete caudal centraScript error: No such module "Unsubst".
Isasicursor[3] I. santacrucensis Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbs An elasmarian ornithopod framless

Sauropods

Sauropods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Nullotitan[3] N. glaciaris Estancia Alta Vista A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle bone A colossosaurian titanosaur framless
Titanosauria Indet.Script error: No such module "Unsubst". IndeterminateScript error: No such module "Unsubst". Seven isolated teethScript error: No such module "Unsubst".

Theropods

Theropods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Kookne[3] K. yeutensis An incomplete right coracoid lacking sternal end and proximal end damaged An ornithuran bird
Maip[4] M. macrothorax La Anita Farm Axis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsal The largest megaraptorid theropod of South America and possibly the largest member of this group

File:Maip macrothorax reconstruction.jpg

Megaraptoridae Indet.Script error: No such module "Unsubst". IndeterminateScript error: No such module "Unsubst". La Anita FarmScript error: No such module "Unsubst". Isolated teethScript error: No such module "Unsubst".
Noasauridae Indet.Script error: No such module "Unsubst". IndeterminateScript error: No such module "Unsubst". Isolated right pedal phalanx IV-2?Script error: No such module "Unsubst".
Unenlagiidae Indet.Script error: No such module "Unsubst". IndeterminateScript error: No such module "Unsubst". An isolated dorsal vertebrae, pedal ungual of digit II and phalanx III-2Script error: No such module "Unsubst".
Yatenavis[5] Y. ieujensis Estancia La Anita. Distal right humerus An enantiornithine bird

Thyreophorans

Thyreophorans of the Chorrillo Formation
Genus Species Locality Material Notes Images
Parankylosauria Indet.[6] Indeterminate Five fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms[7]

Amphibians

Amphibians of the Chorrillo Formation
Genus Species Locality Material Notes Images
Anura Indet. Indeterminate Isolated tibiofibula
Calyptocephalella C. gayi Incomplete left maxilla, incomplete right maxilla, incomplete right ilium, proximal right radioulna, left proximal end of tibiofibula and proximal half of urostyle
Calyptocephalellidae Indet. Indeterminate A distal end of right humerus
Pipoidea Indet. Indeterminate Distal end of right humerus

Fishes

Fishes of the Chorrillo Formation
Genus Species Locality Material Notes Images
Lamniformes Indet. Indeterminate Isolated upper lateral tooth
Teleostei Indet. Indeterminate Three crushing tooth crowns
Vidalamiinae Indet. Indeterminate Isolated tooth A amiid fish

Mammals

Mammals of the Chorrillo Formation
Genus Species Locality Material Notes Images
Magallanodon[8] M. baikashkenke Río de Las Chinas Valley and La Anita Farm Tentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right side A ferugliotheriid gondwanathere
Mammalia Indet. Indeterminate Two isolated caudal vertebrae
Patagorhynchus[9] P. pascuali La Anita Farm Lower right molar attached to a fragment of the dentary A ornithorhynchid monotreme File:Patagorhynchus holotype.png
Patagomaia[10] P. chainko La Anita Farm Partial left acetabulum and ischium A large therian File:Patagomaia chainko.webp

Reptiles

Mosasaurs

Mosasaurs of the Chorrillo Formation
Genus Species Locality Material Notes Images
Mosasauridae Indeterminate

Rhynchocephalian

Rhynchocephalians of the Chorrillo Formation
Genus Species Locality Material Notes Images
Notosphenos[11] N. finisterre A sphenodontid rhynchocephalian

Snakes

Snakes of the Chorrillo Formation
Genus Species Locality Material Notes Images
Rionegrophis R. madtsoioides A partial vertebra. A madtsoiid snake
Serpentes Indeterminate Partial mid-posterior vertebrae

Turtles

Turtles of the Chorrillo Formation
Genus Species Locality Material Notes Images
Chelidae Indet. Indeterminate
Hydromedusinae Indet. Indeterminate Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragment
Meiolaniformes Indet. Indeterminate Distal end of a left humerus

Insects

Insects of the Chorrillo Formation
Genus Species Locality Material Notes Images
Coelolepida Indet.[12] Indeterminate La Anita farm MPM-Pal 21835-3:W34/1, MPM-Pal 21835-23:W55/0, MPM-Pal 21835-22:R52/2,MPM-Pal 21835-20:Z47/3. MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3 A Glossatan Lepidopteran
Diamesinae Indet.[12] "Morphotype 3" La Anita farm MPM-Pal 21835-9:E34/4, MPM-Pal 21835-19: V34/4 A Chironomidae Nematoceran. Similar to the living genus Paraheptagyia
Ephemeroptera Indet.[12] Indeterminate La Anita farm MPM-Pal 21835-9:W37/1 A Mayfly
Lepidoptera Indet.[12] Indeterminate La Anita farm MPM-Pal 21835-10:X30/2, MPM-Pal 21835-7:S27/2
Orthocladiinae Indet.[12] "Morphotype 1" La Anita farm MPM-Pal 21835-24: M31/0, MPM-Pal 21835-3: P46/4 A Chironomidae Nematoceran
"Morphotype 2" La Anita farm MPM-Pal 21835-25: V26/3
"Morphotype 5" La Anita farm MPM-Pal 21835-9: Q58/3 It resembles modern Cricotopus File:Cricotopus P1560901a.jpg
Tanypodinae Indet.[12] "Morphotype 4" La Anita farm MPM-Pal 21835-2:Y40/3 A Chironomidae Nematoceran

Molluscs

Gastropods

Gastropods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Allopeas[13] A. agnolini A achatinid gastropod
Bocourtia[13] B. leonardodavincii A bulimulid gastropod
Lilloiconcha[13] L. novasi A charopid gastropod
Coelostemma[13] C. patagonica A holospirid gastropod
Holospira H. sp.
Physa P. sp.
Pleuroceridae Indet. Indeterminate
Pomacea P. sp.
Potamolithus[13] P. fabiani A tateid gastropod
Stenophysa S. sp.

Plants

Plants of the Chorrillo Formation
Genus Species Locality Material Notes Images
Podocarpoxylon P. dusenii

Algaes

Algaes of the Chorrillo Formation
Genus Species Locality Material Notes Images
Sphaeroplea[14] S. striatocristata

Pollen

Pollen of the Chorrillo Formation
Genus Species Locality Material Notes Images
Acylomurus[14] A. silviae
Baculatisporites B. comaumensis
B. kachaikensis
Ceratosporites C. equalis
Clavatipollenites C. sp.
Clavifera C. triplex
Concavissimisporites C. sp.
Cyatheacidites C. annulatus
Cyathidites C. australis
C. minor
C. rafaeli
Foveosporites F. canalis
Gleicheniidites G. senonicus
Laevigatosporites L. ovatus
Leptolepidites L. sp.
Lusatisporis[14] L. choiols
Microcachryidites M. antarcticus
Peninsulapollis P. gilli
Phyllocladidites P. mawsonii
Podocarpidites cf. P. ellipticus
cf. P. herbstii
P. sp.
Retitriletes R. austroclavatidites
Trichotomosulcites T. subgranulatus
Trilites T. fasolae
cf. T. tuberculiformis
cf. T parvallatus
Tricolpites T. reticulatus
T. sp.
Tuberculatosporites T. parvus
Vitreisporites V. signatus

See also

References

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  1. ↑ Río Leona at Fossilworks.org
  2. ↑ Marenssi et al., 2003, p.406
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Bibliography

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Further reading

  • J. E. Powell. 2003. Revision of South American titanosaurid dinosaurs: palaeobiological, palaeobiogeographical and phylogenetic aspects. Records of the Queen Victoria Museum Launceston 111:1-173