Berothidae

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Template:Short description Template:Automatic taxobox

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings.[1] The family was first named by Anton Handlirsch in 1906.[2] The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions.[3] Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.[4]

Systematics

File:Berothidae Bandipur.jpg
Typical posture in life

A considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or incertae sedis.

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Subfamily Berothimerobiinae
  • Berothimerobius Monserrat & Deretsky, 1999Script error: No such module "Check for unknown parameters".
Subfamily Berothinae
  • Asadeteva Aspöck & Aspöck, 1981Script error: No such module "Check for unknown parameters".
  • Barrowiella Smithers, 1984Script error: No such module "Check for unknown parameters".
  • Berlekrumyia Aspöck & Aspöck, 1988Script error: No such module "Check for unknown parameters".
  • Berotha Walker, 1860Script error: No such module "Check for unknown parameters".
  • †Elektroberotha Makarkin & Ohl 2015Script error: No such module "Check for unknown parameters". Eocene (Lutetian); Baltic amber, northern Europe
  • Isoscelipteron Costa, 1863Script error: No such module "Check for unknown parameters".
  • Lekrugeria Navás, 1929Script error: No such module "Check for unknown parameters".
  • Lomamyia Banks, 1904Script error: No such module "Check for unknown parameters".
  • Nodalla Navás, 1926Script error: No such module "Check for unknown parameters".
  • Podallea Navás, 1936Script error: No such module "Check for unknown parameters".
  • Quasispermophorella Aspöck & Aspöck, 1986Script error: No such module "Check for unknown parameters".
  • Spermophorella Tillyard, 1916Script error: No such module "Check for unknown parameters".
  • Spiroberotha Adams, 1990Script error: No such module "Check for unknown parameters".
  • Stenobiella Tillyard, 1916Script error: No such module "Check for unknown parameters".
  • †Xenoberotha Makarkin 2017Script error: No such module "Check for unknown parameters". Green River Formation, United States, Eocene
Subfamily Cyrenoberothinae[5]
  • †Aggregataberotha Wang et al., 2023Script error: No such module "Check for unknown parameters". (Cenomanian, Burmese amber, Myanmar)
  • †Araripeberotha Martins-Neto & Vulcano 1990Script error: No such module "Check for unknown parameters". Cretaceous (Aptian); Crato Formation, Brazil
  • †Caririberotha Martins-Neto & Vulcano 1990Script error: No such module "Check for unknown parameters". Cretaceous (Aptian); Crato Formation, Brazil
  • Cyrenoberotha MacLeod & Adams, 1968Script error: No such module "Check for unknown parameters".
  • Manselliberotha Aspöck & Aspöck, 1988Script error: No such module "Check for unknown parameters".
  • †Microberotha Archibald & Makarkin 2004Script error: No such module "Check for unknown parameters". (Eocene (Ypresian), Hat Creek amber, Canada)[2]
  • Ormiscocerus Blanchard, 1851Script error: No such module "Check for unknown parameters".
  • †Protoberotha Huang et al., 2019Script error: No such module "Check for unknown parameters". (Cenomanian, Burmese amber, Myanmar)[6]
  • †Sibelliberotha Azar & Nel 2013Script error: No such module "Check for unknown parameters". (Barremian, Lebanese amber)
  • Speleoberotha Machado et al., 2022Script error: No such module "Check for unknown parameters".[5]
Subfamily †Mesithoninae Panfilov, 1980Script error: No such module "Check for unknown parameters".
Subfamily Nosybinae
  • †Banoberotha Whalley 1980Script error: No such module "Check for unknown parameters". Cretaceous (Barremian); Lebanese amber, Lebanon
  • Naizema Navás, 1919Script error: No such module "Check for unknown parameters".
  • Nosybus Navás, 1910Script error: No such module "Check for unknown parameters".
  • Tanzanberotha Aspöck & Hynd, 1995Script error: No such module "Check for unknown parameters".
Subfamily Nyrminae
  • Nyrma Navás, 1933Script error: No such module "Check for unknown parameters".
Subfamily Protobiellinae
  • Austroberothella Aspöck & Aspöck, 1985Script error: No such module "Check for unknown parameters".
  • Protobiella Tillyard, 1923Script error: No such module "Check for unknown parameters".
Subfamily Trichomatinae
  • Trichoberotha Handschin, 1935Script error: No such module "Check for unknown parameters".
  • Trichoma Tillyard, 1916Script error: No such module "Check for unknown parameters".
Subfamily Incertae sedis
  • †Ansoberotha Yang et al., 2019Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Berothone Khramov 2015Script error: No such module "Check for unknown parameters". Callovian/Oxfordian; Karabastau Formation, Kazakhstan
  • †Cantabroberotha Pérez-de la Fuente et al., 2021Script error: No such module "Check for unknown parameters". Cretaceous (Albian); Spanish amber, Spain
  • †Cornoberotha Yang et al., 2019Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Dasyberotha Engel & Grimaldi 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Epimesoberotha Jepson et al., 2012Script error: No such module "Check for unknown parameters". Cretaceous (Berriasian); Lulworth Formation, United Kingdom
  • †Ethiroberotha Engel & Grimaldi 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Haploberotha Engel & Grimaldi 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Iceloberotha Engel & Grimaldi 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Jersiberotha Grimaldi 2000Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar; Cretaceous (Turonian); New Jersey amber, United States
  • †Krokhathone Khramov 2015Script error: No such module "Check for unknown parameters". Jurassic (Callovian/Oxfordian); Karabastau Formation, Kazakhstan
  • †Maculaberotha Yuan et al., 2016Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Magniberotha Yuan et al., 2016Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Nascimberotha Grimaldi 2000Script error: No such module "Check for unknown parameters". Turonian; New Jersey amber, United States
  • †Plesiorobius Klimaszewski & Kevan, 1986Script error: No such module "Check for unknown parameters". Cretaceous (Campanian); Canadian amber, Canada; Campanian; Taymyr amber, Russia; Cenomanian, unknown formation, Russia
  • †Proberotha Krüger, 1923Script error: No such module "Check for unknown parameters". Lutetian, Baltic Amber, northern Europe
  • †Osmyloberotha Khramov 2021Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Sinosmylites Hong 1983Script error: No such module "Check for unknown parameters". Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, Ulaan-Ereg Formation, Mongolia, Late Jurassic (Tithonian)
  • †Systenoberotha Engel & Grimaldi 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Telistoberotha Engel & Grimaldi 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Xiaoberotha Shi et al., 2019Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar

Formerly included taxa

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Subfamily Paraberothinae (now included in Rhachiberothidae)

  • †Acanthoberotha Nakamine et al., 2020Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Alboberotha Nel et al., 2005Script error: No such module "Check for unknown parameters". Cretaceous (Albian); Charentense amber, France
  • †Astioberotha Nakamine et al., 2020Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Chimerhachiberotha Nel et al., 2005Script error: No such module "Check for unknown parameters". Barremian; Lebanese amber, Lebanon
  • †Creagroparaberotha Makarkin, 2015Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †EorhachiberothaEngel, 2004Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Kujiberotha Nakamine & Yamamoto, 2018Script error: No such module "Check for unknown parameters". Cretaceous (Santonian); Kuji amber, Japan
  • †Micromantispa Shi et al., 2015Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Paraberotha Whalley, 1980Script error: No such module "Check for unknown parameters". Barremian; Lebanese amber, Lebanon
  • †Pseudosisyra Makarkin, 1999Script error: No such module "Check for unknown parameters". Aptian; Zaza Formation, Russia
  • †Raptorapax Petrulevicius et al., 2010Script error: No such module "Check for unknown parameters". Barremian; Lebanese amber, Lebanon
  • †Retinoberotha Schlüter, 1978Script error: No such module "Check for unknown parameters". Cenomanian; Bezonnais amber, France,
  • †Rhachibermissa Grimaldi, 2000Script error: No such module "Check for unknown parameters". Turonian; New Jersey amber, United States
  • †Scoloberotha Engel & Grimaldi, 2008Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Spinoberotha Nel et al., 2005Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Stygioberotha Nakamine et al, 2020Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar
  • †Uranoberotha Nakamine et al, 2020Script error: No such module "Check for unknown parameters". Cenomanian; Burmese amber, Myanmar

References

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  1. ↑ Script error: No such module "Citation/CS1".
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  5. ↑ a b Script error: No such module "Citation/CS1".
  6. ↑ Script error: No such module "Citation/CS1".

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