Coconut: Difference between revisions
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{{Short description|Species of plant in the palm family}} | {{Short description|Species of plant in the palm family}} | ||
{{good article}} | |||
{{About|the plant}} | {{About|the plant}} | ||
{{redirect|Coconut tree|other uses|Coconut Tree (disambiguation)}} | {{redirect|Coconut tree|other uses|Coconut Tree (disambiguation)}} | ||
{{pp-semi-indef}} | {{pp-semi-indef}} | ||
{{Use mdy dates|date=July 2014}} | {{Use mdy dates|date=July 2014}} | ||
{{Use American English|date=July 2020}} | {{Use American English|date=July 2020}} | ||
{{Speciesbox | {{Speciesbox | ||
| fossil_range = {{Fossil range|55|0}}<small>Early [[Eocene]] – Recent</small> | |fossil_range = {{Fossil range|55|0}}<small>Early [[Eocene]] – Recent</small> | ||
| image = Cocos_nucifera_-_Köhler–s_Medizinal-Pflanzen-187.jpg | |image = Cocos_nucifera_-_Köhler–s_Medizinal-Pflanzen-187.jpg | ||
|image2 = Coconut 4.jpg | |||
|display_parents = 4 | |||
| image2 = Coconut 4.jpg | |genus = Cocos | ||
|parent_authority = [[Carl Linnaeus|L.]] | |||
|species = nucifera | |||
| display_parents = 4 | |authority = L. | ||
| genus = Cocos | |synonyms = | ||
| parent_authority = [[Carl Linnaeus|L.]] | |||
| species = nucifera | |||
| authority = L. | |||
| synonyms = | |||
* ''Coccus'' {{Au|Mill.}} | * ''Coccus'' {{Au|Mill.}} | ||
* ''Calappa'' {{Au|Steck}} | * ''Calappa'' {{Au|Steck}} | ||
* ''Coccos'' {{Au|Gaertn.}} | * ''Coccos'' {{Au|Gaertn.}} | ||
| synonyms_ref = <ref name="WCSP">{{cite web|url= | |synonyms_ref = <ref name="WCSP">{{cite web |url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:666160-1 |title=''Cocos nucifera'' L. |date=2025 |work=[[World Checklist of Selected Plant Families]] |publisher=[[Royal Botanic Gardens, Kew]] |access-date=22 November 2025}}<!--see 'Synonyms' tab--></ref> | ||
| range_map = Cocos nucifera origin native range map.svg | |range_map = Cocos nucifera origin native range map.svg | ||
| range_map_caption = Possible native range prior to [[crop domestication|domestication]] | |range_map_caption = Possible native range prior to [[crop domestication|domestication]] | ||
}} | }} | ||
The '''coconut | The '''coconut''' ('''''Cocos nucifera''''') is a member of the palm [[family (biology)|family]] ([[Arecaceae]]) and the only living [[species]] of the [[genus]] ''Cocos''.<ref name="WCSP"/> The term "'''coconut'''" (or the archaic "'''cocoanut'''")<ref name="COD">{{cite encyclopedia |title=Coconut |encyclopedia=[[Concise Oxford Dictionary]] |editor=Pearsall, J. |year=1999 |edition=10th |location=Oxford |publisher=[[Clarendon Press]] |isbn=0-19-860287-1}}</ref> can denote the whole '''coconut palm''' [[tree]] or the large hard [[fruit]]. Originally native to [[Central Indo-Pacific]], they are ubiquitous in coastal tropical regions. | ||
The coconut tree provides food, fuel, cosmetics, folk medicine and building materials | The coconut tree provides food, fuel, cosmetics, folk medicine and building materials. The inner flesh of the mature fruit forms a regular part of the diets of many people in the tropics and [[subtropics]]. Coconut [[endosperm]] contains a large quantity of a liquid, "[[coconut water]]". Mature coconuts can be processed for [[Coconut oil|oil]] and [[coconut milk]] from the flesh, [[charcoal]] from the hard shell, and [[coir]] from the fibrous [[husk]]. Dried coconut flesh is called [[copra]], and the oil and milk derived from it are commonly used in cooking and in [[soap]]s and [[cosmetics]]. Sweet coconut sap can be made into drinks or fermented into [[palm wine]] or [[coconut vinegar]]. The hard shells, fibrous husks and long pinnate leaves are used to make a products for [[furnishing]] and decoration. | ||
The coconut has cultural and religious significance | The coconut has cultural and religious significance for [[Austronesian peoples]], appearing in their mythologies, songs, and oral traditions. It has religious significance in [[South Asia]]n cultures, where it is used in [[Hindu]] [[ritual]]s including weddings and worship. | ||
The species evolved in the central Indo-Pacific. It was [[crop domestication|domesticated]] by [[Austronesian peoples]] in [[Island Southeast Asia]] and spread during the [[Neolithic]] via their [[Austronesian expansion|seaborne migrations]] as far east as the [[Pacific Islands]], and as far west as [[Madagascar]]. The species played a critical role in their long sea voyages by providing a portable source of food and water, as well as building materials for [[outrigger boat]]s. Coconuts were spread much later along the coasts of the [[Indian Ocean|Indian]] and [[Atlantic Ocean|Atlantic]] Oceans by [[South Asia]]n, [[Arab people|Arab]], and from the 16th century by European sailors. Based on these introductions, the species can be divided into Pacific and Indo-Atlantic types. The Indo-Atlantic type was introduced to the [[Americas]] during the [[European colonization of the Americas|colonial era]] in the [[Columbian exchange]], while Austronesian sailors appear to have introduced Pacific coconuts to [[Panama]] in [[pre-Columbian]] times. | |||
Trees can grow up to {{convert|30|m|ft|sigfig=1|abbr=off}} tall and can yield up to 75 fruits per year, though fewer than 30 is more typical. | Trees can grow up to {{convert|30|m|ft|sigfig=1|abbr=off}} tall and can yield up to 75 fruits per year, though fewer than 30 is more typical. They are intolerant to cold and prefer copious precipitation and full sunlight. Many insect pests and diseases affect commercial production. In 2023, world production of coconuts was 65 million [[tonne]]s, with 73% of the total produced by [[Indonesia]], [[India]], and the [[Philippines]]. | ||
==Description == | |||
[[File:Coconut tree leave.jpeg|thumb|Coconut palm leaves]] | [[File:Coconut tree leave.jpeg|thumb|Coconut palm leaves]] | ||
''Cocos nucifera'' is a large palm, growing up to {{convert|30|m|ft|abbr=off|sigfig=1}} tall, with [[pinnate]] leaves {{convert|4|-|6|m|ft|0|abbr=on}} long, and pinnae {{convert|60|-|90|cm|ft|0}} long; old leaves break away cleanly, leaving the [[trunk (botany)|trunk]] smooth.<ref name="Pradeepkumar">{{cite book |last1= Pradeepkumar |first1= T. |last2=Sumajyothibhaskar |first2=B. |last3= Satheesan |first3= K.N. |year= 2008 |title= Management of Horticultural Crops |url= https://books.google.com/books?id=VHmokNZXbHUC&pg=PA539 |series= | ''Cocos nucifera'' is a large palm, growing up to {{convert|30|m|ft|abbr=off|sigfig=1}} tall, with [[pinnate]] leaves {{convert|4|-|6|m|ft|0|abbr=on}} long, and pinnae {{convert|60|-|90|cm|ft|0}} long; old leaves break away cleanly, leaving the [[trunk (botany)|trunk]] smooth.<ref name="Pradeepkumar">{{cite book |last1=Pradeepkumar |first1=T. |last2=Sumajyothibhaskar |first2=B. |last3=Satheesan |first3=K. N. |year=2008 |title=Management of Horticultural Crops |url=https://books.google.com/books?id=VHmokNZXbHUC&pg=PA539 |series=Horticulture Science Series |volume=11 |publisher=New India Publishing |pages=539–587 |isbn=978-81-89422-49-3}}</ref> | ||
On fertile soil, a tall coconut palm tree produces around 80 [[fruit]]s per year; new varieties may be able to yield as many as 150 per year.<ref name="Sarian">{{cite news |last=Sarian |first=Zac B. |date=18 August 2010 |title=New coconut yields high |url=http://www.mb.com.ph/articles/272929/new-coconut-yields-high |work=The [[Manila Bulletin]] |archive-url=https://web.archive.org/web/20111119033447/http://mb.com.ph/articles/272929/new-coconut-yields-high |archive-date=19 November 2011 |access-date= 21 April 2011}}</ref> In India, average production is over 8,000 nuts per hectare per year.<ref name="Ravi">{{cite news |last=Ravi |first=Rajesh |date=16 March 2009 |title=Rise in coconut yield, farming area put India on top |url=http://www.financialexpress.com/news/rise-in-coconut-yield-farming-area-put-india-on-top/434818/0 |work=The [[The Financial Express (India)|Financial Express]]|archive-url=https://web.archive.org/web/20130515043307/http://www.financialexpress.com/news/rise-in-coconut-yield-farming-area-put-india-on-top/434818/0 |archive-date=15 May 2013 |access-date=21 April 2011}}</ref> Tall varieties produce their first fruit in 6 to 10 years, and live for 60 to 100 years; dwarf varieties become productive more quickly, but have a shorter lifespan.<ref name="Elzabroek"/> | |||
=== Inflorescence === | |||
[[File:Cocos nucifera (11124377083).jpg|thumb|Inflorescence unfurling]] | |||
The coconut is [[monoecious]], meaning that both male and female [[flower]]s grow on the same tree, in its case in the same inflorescence.<ref name="Thampan 1981">{{cite book |last=Thampan |first=P.K. |year=1981 |title=Handbook on Coconut Palm |publisher=Oxford & IBH Publishing, for Department of Agriculture & Cooperation, Government of India |location=New Delhi, Bombay, and Calcutta |url=https://ia800107.us.archive.org/17/items/in.ernet.dli.2015.201473/2015.201473.Hand-Book.pdf |pages=10–11 |access-date=29 November 2025}}</ref><ref name="Kew Coconut palm">{{cite web |title=Cocos nucifera: Coconut palm |url=https://www.kew.org/plants/coconut-palm |publisher=[[Royal Botanic Gardens, Kew]] |access-date=29 November 2025}}</ref> It is possible that the species in addition occasionally has bisexual flowers.<ref>{{cite book |last=Willmer |first=Pat |url=https://books.google.com/books?id=UVGi6W8QdO4C&pg=PA57 |title=Pollination and Floral Ecology |date=2011-07-25 |publisher=[[Princeton University Press]] |isbn=978-0-691-12861-0 |page=57}}</ref> The female flower is much larger than the male flower.<ref name="Kew Coconut palm"/> Mature trees grow continuously, producing leaves, flowers, and fruit all year round. It takes some 14 months for each flower [[primordium]] to develop into an [[inflorescence]], botanically a [[Spadix (botany)|spadix]] inside a sheathing [[spathe]]. A healthy tree can produce up to 15 inflorescences per year, staggered so that there is always a mature one with others in different stages of development.<ref name="Agriculture Institute 2024">{{cite web |title=Flowering and Fruiting Patterns of Coconut Trees |url=https://agriculture.institute/post-harvest-mgt-value-addition/flowering-fruiting-patterns-coconut-trees/ |publisher=Agriculture Institute |access-date=29 November 2025 |date=27 January 2024 |quote=Under optimal growing conditions, healthy mature trees produce new inflorescences every month, creating a continuous cycle of flowering and fruiting that can yield 12 to 15 spadices per year.}}</ref><ref name="Gordon Jackson 2017">{{cite book |last1=Gordon |first1=André |last2=Jackson |first2=J. |title=Food Safety and Quality Systems in Developing Countries |date=2017 |editor=Gordon, André |volume=2 |chapter=Case study: application of appropriate technologies to improve the quality and safety of coconut water |isbn=978-0-12-801226-0 |pages=185-216}}</ref> It takes 11 months from the opening of the female flowers to the time of harvest.<ref name="Ranasinghe 2015">{{cite journal |last1=Ranasinghe |first1=C.S. |last2=Silva |first2=L.R.S. |last3=Premasiri |first3=R.D.N. |title=Major determinants of fruit set and yield fluctuation in coconut (Cocos nucifera L.) |journal=Journal of the National Science Foundation of Sri Lanka |volume=43 |issue=3 |date=24 September 2015 |doi=10.4038/jnsfsr.v43i3.7955 |page=253 |url=https://jnsfsl.sljol.info/article/10.4038/jnsfsr.v43i3.7955/ |access-date=29 November 2025|doi-access=free }}</ref> Coconut palms are largely cross-[[pollination|pollinated]], although most dwarf varieties are self-pollinating.<ref name="Lutz"/> | |||
=== Fruit === | |||
[[File:Exocarp of coconut.jpg|thumb|upright|Husks]] | |||
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[[Botany|Botanically]], the coconut fruit is a [[drupe]], not a [[Nut (fruit)|true nut]].<ref name="botany">{{cite web |date=December 2014 |title=Coconut botany |url=http://agritech.tnau.ac.in/horticulture/horti_pcrops_coconut_botany.html |access-date=14 December 2017 |website=Agritech Portal |publisher=[[Tamil Nadu Agricultural University]]}}</ref> Like other fruits, it has [[Fruit anatomy|three layers]]: the [[exocarp]], [[mesocarp]], and [[endocarp]]. The exocarp is the glossy outer skin, usually yellow-green to yellow-brown in color. The mesocarp is composed of a [[fiber]], called [[coir]], which has many traditional and commercial uses. | [[Botany|Botanically]], the coconut fruit is a [[drupe]], not a [[Nut (fruit)|true nut]].<ref name="botany">{{cite web |date=December 2014 |title=Coconut botany |url=http://agritech.tnau.ac.in/horticulture/horti_pcrops_coconut_botany.html |access-date=14 December 2017 |website=Agritech Portal |publisher=[[Tamil Nadu Agricultural University]]}}</ref> Like other fruits, it has [[Fruit anatomy|three layers]]: the [[exocarp]], [[mesocarp]], and [[endocarp]]. The exocarp is the glossy outer skin, usually yellow-green to yellow-brown in color. The mesocarp is composed of a [[fiber]], called [[coir]], which has many traditional and commercial uses. The exocarp and the mesocarp make up the "husk" of the coconut, while the endocarp makes up the hard coconut "shell". The endocarp is around {{convert|4|mm|in|abbr=off|frac=8}} thick and has three distinctive [[germination]] pores ([[Micropyle (botany)|micropyle]]s) on the distal end. Two of the pores are plugged (the "eyes"), while one is functional.<ref name="Ledo">{{cite journal |last1=Lédo |first1=Ana da Silva |last2=Passos |first2=Edson Eduardo Melo |last3=Fontes |first3=Humberto Rolemberg |last4=Ferreira |first4=Joana Maria Santos |last5=Talamini |first5=Viviane |last6=Vendrame |first6=Wagner A. |last7=Lédo |first7=Ana da Silva |last8=Passos |first8=Edson Eduardo Melo |last9=Fontes |first9=Humberto Rolemberg |last10=Ferreira |first10=Joana Maria Santos |last11=Talamini |first11=Viviane |last12=Vendrame |first12=Wagner A. |display-authors=5 |date=2019 |title=Advances in Coconut palm propagation |journal=Revista Brasileira de Fruticultura |volume=41 |issue=2 |doi=10.1590/0100-29452019159 |doi-access=free |article-number=e-159}}</ref><ref name="Armstrong">{{cite web |last1=Armstrong |first1=W. P. |title=Edible Palm Fruits |url=https://www2.palomar.edu/users/warmstrong/ecoph10.htm |access-date=20 April 2021 |website=Wayne's Word: An On-Line Textbook of Natural History |publisher=[[Palomar College]] |archive-date=September 2, 2018 |archive-url=https://web.archive.org/web/20180902172839/https://www2.palomar.edu/users/warmstrong/ecoph10.htm }}</ref> | ||
[[File:JfBohol7038naturefvf 39.JPG|thumb|Palm heavy with fruit]] | [[File:JfBohol7038naturefvf 39.JPG|thumb|Palm heavy with fruit]] | ||
The interior of the endocarp is hollow and is lined with a thin brown [[seed coat]] | The interior of the endocarp is hollow and is lined with a thin brown [[seed coat]], some {{convert|0.2|mm|in|abbr=on|frac=64}} thick. The endocarp is initially filled with a liquid [[endosperm]] (the [[coconut water]]). The liquid contains many free [[Cell nucleus|cell nuclei]] (it is [[Multinucleate cell|multinucleate]]) dividing by [[mitosis]], without cell boundaries. As development continues, cellular layers of endosperm deposit along the walls of the endocarp up to {{convert|11|mm|in|abbr=on|frac=8}} thick, starting at the far end. They eventually form the edible solid endosperm ("coconut meat") which hardens over time. The small cylindrical [[plant embryo|embryo]] is embedded in the solid endosperm directly below the functional pore. During germination, the embryo pushes out of the functional pore and forms a [[haustorium]] inside the central cavity. This absorbs the solid endosperm to nourish [[coconut sprout|the seedling]].<ref name="Ledo"/><ref name="duke">{{cite web |date=1983 |title=''Cocos nucifera'' L. (Source: James A. Duke. 1983. Handbook of Energy Crops; unpublished) |url=http://www.hort.purdue.edu/newcrop/duke_energy/Cocos_nucifera.html |url-status=live |archive-url=https://web.archive.org/web/20150603080303/http://www.hort.purdue.edu/newcrop/duke_energy/Cocos_nucifera.html |archive-date=June 3, 2015 |access-date=4 June 2015 |publisher=[[Purdue University]], NewCROP – New Crop Resource}}</ref><ref name="Sugimuma">{{cite journal |last1=Sugimuma |first1=Yukio |last2=Murakami |first2=Taka |date=1990 |title=Structure and Function of the Haustorium in Germinating Coconut Palm Seed |url=https://www.jircas.go.jp/sites/default/files/publication/jarq/24-1-001-014_0.pdf |journal=JARQ |volume=24 |pages=1–14}}</ref> | ||
The fruits have two distinctive forms. Wild ''niu kafa'' coconuts feature an elongated triangular fruit with a thicker husk and a smaller amount of endosperm. These make the fruits more buoyant, ideal for ocean dispersal. Domesticated ''niu vai'' Pacific coconuts are rounded in shape with a thinner husk, more endosperm, and more [[coconut water]].<ref name="Lebrun"/><ref name="Shukla"/><ref name="Lutz"/> | |||
Domesticated Pacific coconuts | |||
A full-sized | {| class="wikitable floatright" | ||
|+ Composition of a<br/>Bido coconut<ref name="Novarianto 2022"/> | |||
|- | |||
! colspan=2 | Component | |||
! colspan=2 | Mass/kg | |||
|- | |||
| colspan=3 | Husk || 2.033 | |||
|- | |||
| colspan=3 | Nut || 1.125 | |||
|- | |||
| rowspan=3 | of which || Shell || 0.359 | |||
|- | |||
| Juice || 0.492 | |||
|- | |||
| Meat || 0.477 | |||
|- | |||
| colspan=3 | Total || 3.158 | |||
|} | |||
A full-sized fruit weighs about {{convert|1.4|kg|lboz|abbr=off}} depending on variety.<ref name="Paull"/><ref name="postharvest"/> The 21st century Bido variety from Indonesia averages {{convert|3.158|kg|lboz|abbr=off}} per fruit.<ref name="Novarianto 2022">{{cite journal |last1=Novarianto |first1=H. |last2=Mawardi |first2=S. |last3=Tulalo |first3=M. A. |title=The Bido variety is an essential genetic material for coconut breeding |journal=IOP Conference Series: Earth and Environmental Science |volume=974 |issue=1 |date=1 January 2022 |doi=10.1088/1755-1315/974/1/012056 |article-number=012056 |url=https://iopscience.iop.org/article/10.1088/1755-1315/974/1/012056 |access-date=29 November 2025|doi-access=free }}</ref> | |||
Coconuts are exported without husks; de-husked nuts from [[Côte d'Ivoire]] average around 575 grams, while de-husked nuts from the [[Dominican Republic]] average nearly 700 grams.<ref>{{cite news |last1=Gerbaud |first1=Pierre |last2=Audibert |first2=Olivier |last3=Bourdeix |first3=Roland |title=Close-up: Coconut |url=https://agritrop.cirad.fr/562361/1/ID562361.pdf |access-date=29 November 2025 |work=FRuiTROP |issue=193 |publisher=CIRAD |date=October 2011}}</ref> | |||
Coconuts sold domestically in coconut-producing countries are typically not de-husked. Immature coconuts (6 to 8 months from flowering) are sold for coconut water and softer jelly-like coconut meat (known as "green coconuts", "young coconuts", or "water coconuts"), where the original coloration of the fruit is more pleasing.<ref name="Paull">{{cite book |last=Paull |first=Robert E. |url=https://www.ctahr.hawaii.edu/oc/freepubs/pdf/F_N-45.pdf |title=Coconut: Postharvest Quality-Maintenance Guidelines |last2=Ketsa |first2=Saichol |date=March 2015 |publisher=College of Tropical Agriculture and Human Resources, [[University of Hawai{{okina}}i at Mānoa]]}}</ref><ref name="postharvest">{{cite book |url=http://agriculture.gov.gy/wp-content/uploads/2016/01/Coconut.pdf |title=Coconut: Postharvest Care and Market Preparation |date=May 2004 |publisher=Ministry of Fisheries, Crops and Livestock, New Guyana Marketing Corporation, National Agricultural Research Institute |series=Technical Bulletin No. 27}}</ref> | |||
Whole mature coconuts (11 to 13 months from flowering) sold for export, however, typically have the husk removed to reduce weight and volume for transport. This results in the naked coconut "shell" with three pores more familiar in countries where coconuts are not grown locally. De-husked | Whole mature coconuts (11 to 13 months from flowering) sold for export, however, typically have the husk removed to reduce weight and volume for transport. This results in the naked coconut "shell" with three pores, remnants of the three carpels of the flower, more familiar in countries where coconuts are not grown locally. De-husked coconuts are easier for consumers to open, but have a shorter postharvest storage life of around two to three weeks at temperatures of {{convert|12|to|15|°C|°F|abbr=on}} or up to 2 months at {{convert|0|to|1.5|°C|°F|abbr=on}}. In comparison, mature coconuts with a husk can be stored for three to five months at room temperature.<ref name="Paull"/><ref name="postharvest"/> | ||
===Roots=== | === Roots === | ||
The [[palm tree]] has neither a [[taproot]] nor [[root hair]]s, but a [[fibrous root system]].<ref name="Thampan 1981"/> This consists of many thin roots that grow outward from the plant near the surface. Only a few penetrate deep into the soil for stability. This is known as a fibrous or adventitious root system, and is a characteristic of grass species. 2,000–4,000 [[adventitious root]]s may grow, each about {{convert|1|cm|in|frac=2|abbr=on}} in diameter. Decayed roots are replaced regularly as the tree grows new ones.<ref name="agro">{{cite book |date=April 2006 |editor=Elevitch, C. R. |chapter=''Cocos nucifera'' (coconut), version 2.1 |url=http://www.agroforestry.net/tti/Cocos-coconut.pdf |url-status=live |archive-url=https://web.archive.org/web/20131020005217/http://agroforestry.net/tti/Cocos-coconut.pdf |archive-date=October 20, 2013 |access-date=22 December 2016 |title=Species Profiles for Pacific Island Agroforestry |publisher=Permanent Agriculture Resources |location=Hōlualoa, Hawaii}}</ref> | |||
[[ | |||
== Taxonomy == | == Taxonomy == | ||
=== Taxonomic history === | |||
The Swedish botanist and taxonomist [[Carl Linnaeus]] formally described the species ''Cocos nucifera'' in his book ''[[Species Plantarum]]'' in 1753.<ref>{{cite book |last=Linnaeus |first=Carl |author-link=Carl Linnaeus |title=[[Species Plantarum]] |year=1753 |page=1188}}</ref> The name is accepted by botanists.<ref name="WCSP"/> In 1768, in his book ''[[The Gardeners Dictionary]]'', the English botanist [[Philip Miller]] redescribed the plant as ''Palma cocos'', a name treated as a synonym.<ref>{{cite book |last=Miller |first=Philip |author-link=Philip Miller |title=[[The Gardeners Dictionary]] |edition=8th |volume=2 |year=1768}}</ref><ref name="WCSP"/> In 1891, the German botanist [[Otto Kuntze]] gave it the name ''Calappa nucifera'' in his ''Revisio Generum Plantarum'', also treated as a synonym.<ref>{{cite book |last=Kuntze |first=Otto |author-link=Otto Kuntze |title=[[Revisio Generum Plantarum]] |volume=2 |page=982 |year=1891}}</ref><ref name="WCSP"/> | |||
=== Etymology === | === Etymology === | ||
[[ | The [[Generic name (biology)|generic name]] ''Cocos'', and the common name, is derived from the 16th-century [[Portuguese language|Portuguese]] word ''[[Coco (folklore)|coco]]'', meaning 'head' or 'skull' after the three indentations on the coconut shell that give an impression of a face.<ref>{{cite book |last=Dalgado |first=Sebastião |url=https://archive.org/details/glossriolusoas00dalguoft/page/n363/mode/2up |title=Glossário luso-asiático |publisher=Coimbra, Imprensa da Universidade |year=1919 |volume=1 |page=291}}</ref><ref>{{cite web |url=https://www.etymonline.com/word/coco |title=coco |website=Online Etymology Dictionary |access-date=3 May 2020}}</ref><ref>{{cite web |url=https://www.etymonline.com/word/coconut |title=coconut |website=Online Etymology Dictionary |access-date=3 May 2020 }}</ref><ref name="Losada">{{cite book |last=Losada |first=Fernando Díez |year=2004 |title=La tribuna del idioma |url=https://books.google.com/books?id=RxGv6tMX2QcC&pg=PT481 |language=es |publisher=Editorial Tecnologica de CR |page=481 |isbn=978-9977-66-161-2}}</ref> This apparently came from encounters in 1521 by [[Portuguese people|Portuguese]] and [[Spanish people|Spanish]] explorers with [[Pacific Islander]]s, when the coconut shell reminded them of [[ghost]]s in Portuguese folklore called ''[[Coco (folklore)|coco]]'' or ''côca''.<ref name="Losada"/><ref>{{cite book |last=Figueiredo |first=Cândido |year=1940 |title=Pequeno Dicionário da Lingua Portuguesa |language=pt |location=Lisbon |publisher=Livraria Bertrand}}</ref> In the West, the fruit was originally called ''nux indica'', a name used by [[Marco Polo]] in 1280 while in [[Sumatra]]. His term is a translation from the Arabic of the time, where it was called جوز هندي ''jawz hindī'', "Indian nut".<ref name="Elzabroek">{{cite book |last=Elzebroek |first=A. T. G. |url=https://books.google.com/books?id=YvU1XnUVxFQC&pg=PA186 |title=Guide to Cultivated Plants |date=2008 |publisher=[[CAB International|CABI]] |isbn=978-1-84593-356-2 |pages=186–192}}</ref> ''Thenga'', its [[Tamil language|Tamil]]/[[Malayalam]] name, was used in the detailed description of coconut found in ''[[Itinerario]]'' by [[Ludovico di Varthema]] published in 1510 and in the later ''[[Hortus Indicus Malabaricus]]''.<ref name="Grimwood1">[[#Grimwood|Grimwood]], p. 1.</ref> | ||
The [[specific name (botany)|specific name]] ''nucifera'' | The [[specific name (botany)|specific name]] ''nucifera'' means "nut-bearing", from the [[Latin]] words ''nux'' (nut) and ''fera'' (bearing).<ref>{{cite web |title=National Flower – Nelumbo nucifera |url=http://bsienvis.nic.in/files/National%20Flower_Nelumbo%20nucifera_26.9.14.pdf |access-date=19 February 2021 |publisher=ENVIS Resource Partner on Biodiversity}}</ref> | ||
== | == Origins == | ||
=== Fossil history === | |||
[[File:Miocene coconut.jpg|thumb|Fossil ''Cocos zeylandica'' from the [[Miocene]] of [[New Zealand]], approximately the size of a [[strawberry]] at {{convert|3.5|cm|in|frac=4|abbr=on}} long|alt=Small blackened fossil]] | |||
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{{ | The vast majority of ''Cocos''-like fossils have been recovered from only two regions in the world: [[New Zealand]] and west-central [[India]]. However, ''Cocos''-like fossils are still putative, as they are difficult to identify.<ref name="Nayar2016">{{cite book |last=Nayar |first=N. Madhavan |url=https://books.google.com/books?id=pWEuDAAAQBAJ |title=The Coconut: Phylogeny, Origins, and Spread |publisher=[[Academic Press]] |year=2016 |isbn=978-0-12-809779-3 |pages=51–66}}</ref> The earliest ''Cocos''-like fossil to be found was ''[[Cocos zeylandica|C. zeylandica]]'', a fossil species with small fruits, around {{convert|3.5|cm|in|frac=4|abbr=on}} × {{convert|1.3|to|2.5|cm|in|frac=8|abbr=on}} in size, from the [[Miocene]] (~23 to 5.3 million years ago) of [[New Zealand]]<!--in 1926-->. Since then, numerous other fossils of similar fruits of uncertain affinity have been found in New Zealand from the [[Eocene]], [[Oligocene]], and possibly the [[Holocene]].<ref name="Nayar2016"/><ref name="Conran2015">{{cite journal|publisher=[[Linnean Society]]|last1=Conran |first1=John G. |last2=Bannister |first2=Jennifer M. |last3=Lee |first3=Daphne E. |last4=Carpenter |first4=Raymond J. |last5=Kennedy |first5=Elizabeth M. |last6=Reichgelt |first6=Tammo |last7=Fordyce |first7=R. Ewan |display-authors=5 |date=2015 |title=An update of monocot macrofossil data from New Zealand and Australia |journal=[[Botanical Journal of the Linnean Society]]|volume=178 |issue=3 |pages=394–420 |doi=10.1111/boj.12284 |doi-access=free}}</ref> In the [[Deccan Traps]] of west-central India, numerous fossils of ''Cocos''-like fruits, leaves, and stems have been found. They include [[morphotaxa]] like ''Palmoxylon sundaran'', ''Palmoxylon insignae'', and ''Palmocarpon cocoides''. ''Cocos''-like fossils of fruits include ''[[Cocos intertrappeansis]]'', ''Cocos pantii'', and ''Cocos sahnii''. Some have been tentatively identified as modern ''C. nucifera''. These include two specimens named ''C. palaeonucifera'' and ''C. binoriensis'', both dated by their authors to the [[Maastrichtian]]–[[Danian]] of the early [[Tertiary (period)|Tertiary]] (70 to 62 million years ago). ''C. binoriensis'' has been claimed to be the earliest known fossil of ''C. nucifera''.<ref name="Srivastava2014">{{cite journal |last1=Srivastava |first1=Rashmi |last2=Srivastava |first2=Gaurav |date=2014 |title=Fossil fruit of ''Cocos'' L. (Arecaceae) from Maastrichtian-Danian sediments of central India and its phytogeographical significance |url=https://www.researchgate.net/publication/261107861 |journal=Acta Palaeobotanica |volume=54 |issue=1 |pages=67–75 |doi=10.2478/acpa-2014-0003 |doi-access=free}}</ref><ref name="Nayar2016"/><ref name="Singh2016">{{cite journal |last1=Singh |first1=Hukam |last2=Shukla |first2=Anumeha |last3=Mehrotra |first3=R. C. |date=2016 |title=A Fossil Coconut Fruit from the Early Eocene of Gujarat |url=http://www.geosocindia.org/index.php/jgsi/article/view/88633 |journal=[[Journal of the Geological Society of India]] |volume=87 |issue=3 |pages=268–270 |doi=10.1007/s12594-016-0394-9 |bibcode=2016JGSI...87..268S |access-date=10 January 2019 |s2cid=131318482 |url-access=subscription }}</ref> | ||
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Only two other regions have reported ''Cocos''-like fossils, namely [[Australia]] and [[Colombia]]. In Australia, a ''Cocos''-like fossil fruit, measuring {{convert|10|x|9.5|cm|in|frac=8|abbr=on}}, was recovered from the Chinchilla Sand Formation dated to the latest [[Pliocene]] or basal [[Pleistocene]]. Rigby (1995) assigned them to modern ''Cocos nucifera'' based on its size.<ref name="Srivastava2014"/><ref name="Nayar2016"/> In Colombia, a single ''Cocos''-like fruit was recovered from the [[Paleocene|middle to late Paleocene]] [[Cerrejón Formation]]. The fruit, however, was compacted in the fossilization process and it was not possible to determine if it had the diagnostic three pores that characterize members of the tribe [[Cocoseae]]. Nevertheless, one study assigned it to ''Cocos'' based on the size and the ridged shape of the fruit.<ref name="Gomez-Navarro2009">{{cite journal |last1=Gomez-Navarro |first1=Carolina |last2=Jaramillo |first2=Carlos |last3=Herrera |first3=Fabiany |last4=Wing |first4=Scott L. |last5=Callejas |first5=Ricardo |date=2009 |title=Palms (Arecaceae) from a Paleocene rainforest of northern Colombia |journal=[[American Journal of Botany]] |volume=96 |issue=7 |pages=1300–1312 |doi=10.3732/ajb.0800378 |pmid=21628279}}</ref> | |||
=== Phylogeny === | |||
A 2016 [[Molecular phylogenetics|molecular phylogenomic]] analysis of the [[Arecaceae|palms]] places the genus ''Cocos'' among the tribe [[Cocoseae]]:<ref>{{cite book |last=Gunn |first=Bee Fong |title=Phylogenomics of the coconut (Cocos nucifera L.) |year=2016 |publisher=[[Australian National University]] (PhD thesis) |url=https://openresearch-repository.anu.edu.au/bitstreams/98d7d3c7-600e-4ead-963e-337362a11b0e/download}}</ref> | |||
=== | {{clade | ||
|label1=[[Cocoseae]] | |||
|1={{clade | |||
|1=''[[Bactris]]'' spiny palms of C. and S. America | |||
|2={{clade | |||
|1=''[[Elaeis oleifera]]'' (American oil palm) of C. and S. America | |||
|2={{clade | |||
|1={{clade | |||
|1=''[[Beccariophoenix madagascariensis]]'' (coastal beccariophoenix) of Madagascar | |||
|2={{clade | |||
|1=''[[Voanioala gerardii]]'' (forest coconut) of Madagascar | |||
|2=''[[Jubaeopsis]]'' (Pondoland palm) of S. Africa | |||
}} | |||
}} | |||
|2={{clade | |||
|1={{clade | |||
|1=''[[Jubaea chilensis]]'' (Chilean wine palm) of central Chile | |||
|2=''[[Butia]]'' (feather palms) of S. America | |||
}} | |||
|2={{clade | |||
|1={{clade | |||
|1=''[[Attalea (plant)|Attalea]]'' non-spiny palms of C. and S. America | |||
|2={{clade | |||
|1=''[[Parajubaea]]'' palms of the Northern Andes | |||
|2=''[[Allagoptera]]'' monoecious palms of S. America | |||
}} | |||
}} | |||
|2={{clade | |||
|1='''''Cocos nucifera''''' ('''coconut palm''') | |||
|2=''[[Syagrus (plant)|Syagrus]]'' palms of S. America | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
=== Human dispersal === | |||
{{main|Origin and dispersal of the coconut}} | |||
[[File:Wa'a kaulua.jpg|thumb|left|[[Catamaran]]s allowed [[Austronesian peoples|Austronesians]] to colonize the islands of the [[Indo-Pacific]] and introduce coconuts as they migrated.<ref name="Mahdi1999">{{cite book |last=Mahdi |first=Waruno |title=Archaeology and Language III: Artefacts languages, and texts |publisher=[[Routledge]] |year=1999 |isbn=978-0-415-10054-0 |editor-last=Blench |editor-first=Roger |series=One World Archaeology |volume=34 |pages=144–179 |chapter=The Dispersal of Austronesian boat forms in the Indian Ocean |editor-last2=Spriggs |editor-first2=Matthew}}</ref><ref name="Doran1981">{{cite book |last1=Doran |first1=Edwin B. |title=Wangka: Austronesian Canoe Origins |date=1981 |publisher=[[Texas A&M University Press]] |isbn=978-0-89096-107-0}}</ref><ref name="Johns">{{cite journal |last1=Johns |first1=D. A. |last2=Irwin |first2=G. J. |last3=Sung |first3=Y. K. |date=29 September 2014 |title=An early sophisticated East Polynesian voyaging canoe discovered on New Zealand's coast |journal=[[PNAS]] |volume=111 |issue=41 |pages=14728–14733 |bibcode=2014PNAS..11114728J |doi=10.1073/pnas.1408491111 |pmc=4205625 |pmid=25267657 |doi-access=free}}</ref>]] | |||
=== | Genetic studies identify the coconut's center of origin as the [[Central Indo-Pacific]], where it has its greatest genetic diversity.<ref name="Perera">{{cite book |last1=Vollmann |first1=Johann |url=https://books.google.com/books?id=Y_heb_lB3qoC&pg=PA370 |title=Oil Crops |last2=Rajcan |first2=Istvan |date=2009-09-18 |publisher=Springer Science & Business Media |isbn=978-0-387-77594-4 |pages=370–372}}</ref><ref name="agro"/><ref name="Baudouin2008">{{cite journal |last1=Baudouin |first1=Luc |last2=Lebrun |first2=Patricia |date=26 July 2008 |title=Coconut (''Cocos nucifera'' L.) DNA studies support the hypothesis of an ancient Austronesian migration from Southeast Asia to America |journal=Genetic Resources and Crop Evolution |volume=56 |issue=2 |pages=257–262 |doi=10.1007/s10722-008-9362-6 |s2cid=19529408}}</ref><ref name="Gunn2011">{{cite journal |last1=Gunn |first1=Bee F. |last2=Baudouin |first2=Luc |last3=Olsen |first3=Kenneth M. |last4=Ingvarsson |first4=Pär K. |date=22 June 2011 |title=Independent Origins of Cultivated Coconut (''Cocos nucifera'' L.) in the Old World Tropics |journal=[[PLOS One]] |volume=6 |issue=6 |article-number=e21143 |bibcode=2011PLoSO...621143G |doi=10.1371/journal.pone.0021143 |pmc=3120816 |pmid=21731660 |doi-access=free}}</ref> Its cultivation and spread was closely tied to migrations of the [[Austronesian peoples]] who [[canoe plants|carried coconuts]] to the islands they settled.<ref name="Baudouin2008"/><ref name="Gunn2011"/><ref name="Crowther2016">{{cite journal |last1=Crowther |first1=Alison |last2=Lucas |first2=Leilani |last3=Helm |first3=Richard |last4=Horton |first4=Mark |last5=Shipton |first5=Ceri |last6=Wright |first6=Henry T. |last7=Walshaw |first7=Sarah |last8=Pawlowicz |first8=Matthew |last9=Radimilahy |first9=Chantal |last10=Douka |first10=Katerina |last11=Picornell-Gelabert |first11=Llorenç |last12=Fuller |first12=Dorian Q. |last13=Boivin |first13=Nicole L. |display-authors=5 |date=14 June 2016 |title=Ancient crops provide first archaeological signature of the westward Austronesian expansion |journal=PNAS |volume=113 |issue=24 |pages=6635–6640 |doi=10.1073/pnas.1522714113 |pmc=4914162 |pmid=27247383 |doi-access=free |bibcode=2016PNAS..113.6635C }}</ref><ref name="Brouwers2011">{{cite web |last1=Brouwers |first1=Lucas |date=1 August 2011 |title=Coconuts: not indigenous, but quite at home nevertheless |url=https://blogs.scientificamerican.com/thoughtomics/httpblogsscientificamericancomthoughtomics20110801coconuts-not-indigenous-but-quite-at-home-nevertheless/ |access-date=10 January 2019 |website=Scientific American}}</ref> Linguistic, archaeological, and genetic evidence all points to domestication of Pacific coconuts by Austronesians in Southeast Asia during the [[Austronesian expansion]] (c. 3000 to 1500 BCE).<ref name="Lew"/><ref name="Gunn2011"/><ref name="Baudouin2008"/><ref name="scid"/> Drift models based on wind and ocean currents show that coconuts could not have drifted across the Pacific unaided,<ref name=Ward/> implying that dispersal was human-assisted.<ref name="coco">{{cite journal |last=Gunn |first=Bee |last2=Baudouin |first2=Luc |last3=Olsen |first3=Kenneth M. |title=Independent Origins of Cultivated Coconut (''Cocos nucifera'' L.) in the Old World Tropics |journal=[[PLOS One]] |year=2011 |volume=6 |issue=6 |article-number=e21143 |doi=10.1371/journal.pone.0021143 |pmid=21731660 |pmc=3120816 |bibcode=2011PLoSO...621143G |doi-access=free}}</ref> | ||
{{multiple image | {{multiple image | ||
|direction = vertical | |||
|width = 300 | |||
|image2 = Historical Introduction of Coconuts (Gunn, Baudouin,& Olsen, 2011).png | |||
|caption2 = Inferred historical introduction of coconuts from the original centers of diversity in the [[Indian subcontinent]] and [[Island Southeast Asia]]<ref name="Gunn2011"/><ref name="Lutz">{{cite web |last1=Lutz |first1=Diana |date=24 June 2011|title=Deep history of coconuts decoded |url=https://source.wustl.edu/2011/06/deep-history-of-coconuts-decoded/ |website=The Source |access-date=10 January 2019}}</ref><ref name="Brouwers2011"/> | |||
|image1 = Geographical distributions of Indo-Atlantic and Pacific coconut subpopulations.png | |||
|caption1 = Geographical distributions of Indo-Atlantic and Pacific coconut subpopulations and their genetic composition<ref name="Gunn2011"/> | |||
}} | }} | ||
Coconuts are divided into two subpopulations, a Pacific group from [[Island Southeast Asia]] and an Indo-Atlantic group from the south of the [[Indian subcontinent]]. The Pacific group is clearly domesticated, with dwarf habit, self-pollination, and ''niu vai'' fruits with large endosperm-to-husk ratios. The distribution of Pacific coconuts corresponds to regions settled by Austronesian voyagers, especially [[Madagascar]]. The island's coconuts show genetic admixture between the two subpopulations, indicating that Pacific coconuts interbred with Indo-Atlantic coconuts there.<ref name="Gunn2011"/><ref name="Crowther2016"/> Although archaeological remains from 1000 to 500 BCE suggest that Indo-Atlantic coconuts were later independently cultivated by [[Dravidian peoples]], only Pacific coconuts show clear domestication traits like dwarf habits, self-pollination, and rounded fruits. Indo-Atlantic coconuts, in contrast, have the ancestral traits of tall habits and elongated triangular fruits.<ref name="Lew"/><ref name="Gunn2011"/><ref name="Baudouin2008"/><ref name="scid">{{cite news |title=Deep history of coconuts decoded: Origins of cultivation, ancient trade routes, and colonization of the Americas |url=https://www.sciencedaily.com/releases/2011/06/110624142037.htm |access-date=22 April 2021 |work=[[ScienceDaily]] |publisher=[[Washington University in St. Louis]] |date=24 June 2011}}</ref> | |||
Genetic studies have confirmed [[pre-Columbian]] populations of coconuts in [[Panama]]. However, it is not native and displays a genetic bottleneck resulting from a [[founder effect]]. Coconuts in the Americas are most closely related to those in the [[Philippines]], indicating that the coconuts were not introduced naturally, such as by sea currents, but by early Austronesian sailors to the Americas from at least 4250 BCE.<ref name="Baudouin2008"/><ref name="Brouwers2011"/><ref name=Ward>{{cite journal |last1=Ward |first1=R. G. |last2=Brookfield |first2=M. |year=1992 |title=Special Paper: the dispersal of the coconut: did it float or was it carried to Panama? |journal=[[Journal of Biogeography]] |volume=19 |issue=5 |pages=467–480 |doi=10.2307/2845766 |jstor=2845766 |bibcode=1992JBiog..19..467W }}</ref> During the [[Early modern period|colonial era]], Pacific coconuts were further introduced to [[Mexico]] from the [[Spanish East Indies]] via the [[Manila galleon]]s, starting in the 16th century. In contrast, Indo-Atlantic coconuts were spread by Arab and Persian traders into the [[East Africa]]n coast. Indo-Atlantic coconuts were introduced into the [[Atlantic Ocean]] by [[Portuguese people|Portuguese]] ships from [[Portuguese India|colonies in India]] and [[Sri Lanka]], again starting in the 16th century: first to coastal [[West Africa]], and then to the [[Caribbean]] and [[Brazil]].<ref name="Gunn2011"/> | |||
=== Domestication === | |||
Coconuts can be broadly divided into two fruit types – the ancestral ''niu kafa'' form with a thick-husked, angular fruit, and the ''niu vai'' form with a thin-husked, spherical fruit with a higher proportion of [[endosperm]]. The terms are [[Samoan language|Samoan]].<ref name="Gunn2011"/><ref name=Lebrun/><ref name=Harries/> | |||
The ''niu | The ''niu kafa'' form is the wild ancestral type, with thick husks to protect the seed, and an angular, highly ridged shape to promote buoyancy during ocean dispersal. It is the dominant form in the Indo-Atlantic coconuts.<ref name=Lebrun/><ref name="Gunn2011"/> However, they may have been selected to some extent for thicker husks for [[coir]] production, which was important in Austronesian material culture as a source for cordage in building houses and boats.<ref name="Lew"/> | ||
<gallery class=center mode=nolines widths=180 heights=180 caption="Two major fruit types"> | |||
File:Cocos nucifera (coconut) 5 (38507429165).jpg|The ''niu kafa'' form of wild and Indo-Atlantic coconuts, possibly selected for more [[coir]] for houses and boats | |||
File:Cocos nucifera (coconut) 2 (39384972311).jpg|The ''niu vai'' form of domesticated Pacific coconuts, [[Selective breeding|selected]] for more meat and [[coconut water|water]], and less husk | |||
</gallery> | |||
The | The ''niu vai'' form is the domesticated form dominant in Pacific coconuts. They were [[Selective breeding|artificially selected]] by Austronesians for their larger endosperm-to-husk ratio and higher coconut water content, making them more useful as food and water reserves for sea voyages. The decreased buoyancy and increased fragility of this spherical, thin-husked fruit did not matter for a species that had started to be dispersed by humans and grown in plantations.<ref name=Lebrun>{{cite journal |last1=Lebrun |first1=P. |last2=Seguin |first2=M. |last3=Grivet |first3=L. |last4=Baudouin |first4=L. |year=1998 |title=Genetic diversity in coconut (''Cocos nucifera'' L.) revealed by restriction fragment length polymorphism (RFLP) markers |doi=10.1023/a:1018323721803 |journal=Euphytica |volume=101 |pages=103–108 |s2cid=19445166}}</ref><ref name=Shukla>{{cite journal |last1=Shukla |first1=A. |last2=Mehrotra |first2=R. C. |last3=Guleria |first3=J. S. |year=2012 |title=Cocos sahnii Kaul: A ''Cocos nucifera'' L.-like fruit from the Early Eocene rainforest of Rajasthan, western India |journal=Journal of Biosciences |volume=37 |issue=4 |pages=769–776 |doi=10.1007/s12038-012-9233-3 |pmid=22922201 |s2cid=14229182 }}</ref> ''Niu vai'' endocarp fragments have been recovered in archaeological sites in the [[St. Matthias Islands]] of the [[Bismarck Archipelago]]. The fragments are dated to approximately 1000 BCE, suggesting that cultivation and artificial selection of coconuts were already practiced by the Austronesian [[Lapita people]].<ref name="Lew"/> | ||
Coconuts can be broadly divided into two general types based on habit: the "Tall" (var. ''typica'') and "Dwarf" (var. ''nana'') varieties.<ref name=Santos>{{cite web |url=http://www.bioversityinternational.org/fileadmin/bioversity/publications/Web_version/108/ch02.htm#Chapter+1+BOTANY+OF+THE+COCONUT+PALM |archive-url=https://web.archive.org/web/20131212200527/http://www.bioversityinternational.org/fileadmin/bioversity/publications/Web_version/108/ch02.htm |title=Chapter 1: Botany of the Coconut Palm |archive-date=December 12, 2013|website=www.bioversityinternational.org}}</ref> The two groups are genetically distinct, with the dwarf variety showing a greater degree of artificial selection for ornamental traits and for early germination and fruiting.<ref name=Harries>{{cite journal |last1=Harries |first1=H. C. |year=1978 |title=The evolution, dissemination and classification of ''Cocos nucifera'' L. |journal=[[The Botanical Review]] |volume=44 |issue=3 |pages=265–319 |doi=10.1007/bf02957852|bibcode =1978BotRv..44..265H |s2cid=26636137 }}</ref><ref name=Huang>{{cite journal |last1=Huang |first1=Y.-Y. |last2=Matzke |first2=A. J. M. |last3=Matzke |first3=M. |year=2013 |title =Complete sequence and comparative analysis of the chloroplast genome of coconut palm (''Cocos nucifera'') |journal=[[PLOS One]] |volume=8 |issue=8 |article-number=e74736 |doi=10.1371/journal.pone.0074736 |pmid=24023703 |pmc=3758300 |bibcode =2013PLoSO...874736H |doi-access=free }}</ref> The tall variety is [[outcrossing]] while dwarf palms are [[Autogamy|self-pollinating]], which has led to a much greater degree of [[genetic diversity]] within the tall group.<ref name="Rivera">{{cite journal |last1=Rivera |first1=R. |last2=Edwards |first2=K. J. |last3=Barker |first3=J. H. |last4=Arnold |first4=G. M. |last5=Ayad |first5=G. |last6=Hodgkin |first6=T. |last7=Karp |first7=A. |title=Isolation and characterization of polymorphic microsatellites in ''Cocos nucifera'' L |journal =Genome |volume =42 |issue =4 |pages =668–675 |year=1999 |pmid =10464790 |doi=10.1139/g98-170}}</ref> | |||
The dwarf coconut cultivars are fully domesticated, unlike the more diverse tall cultivars.<ref name="Bourdeix"/><ref name="Rivera"/> Dwarf coconuts share three genetic markers out of thirteen (rare in tall cultivars), making it likely that they originate from a single domesticated population. Philippine and Malayan dwarf coconuts diverged early into two distinct types. They usually remain genetically isolated when introduced to new regions. Numerous other dwarf cultivars developed after such introductions, hybridizing with tall cultivars. The origin of dwarf varieties is [[Southeast Asia]], which contain the tall cultivars genetically closest to dwarf coconuts.<ref name="Gunn2011"/><ref name="lebrun2013"/><ref name="Bourdeix">{{cite book |last1=Bourdeix |first1=R. |last2=Prades |first2=A. |title=A Global Strategy for the conservation and use of Coconut Genetic Resources 2018–2028 |date=2017 |publisher=[[Bioversity International]] |isbn=978-92-9043-984-4 |url=https://www.bioversityinternational.org/e-library/publications/detail/a-global-strategy-for-the-conservation-and-use-of-coconut-genetic-resources-2018-2028/}}</ref><ref name=Rivera/> | |||
=== | [[Genome sequencing]] of tall and dwarf varieties reveals that they diverged 2 to 8 million years ago and that the dwarf variety arose through alterations in genes for the metabolism of the plant hormone [[gibberellin]].<ref>{{cite journal |last1=Wang |first1=Shouchuang |last2=Xiao |first2=Yong |last3=Zhou |first3=Zhi-Wei |last4=Yuan |first4=Jiaqing |last5=Guo |first5=Hao |last6=Yang |first6=Zhuang |last7=Yang |first7=Jun |last8=Sun |first8=Pengchuan |last9=Sun |first9=Lisong |last10=Deng |first10=Yuan |last11=Xie |first11=Wen-Zhao |last12=Song |first12=Jia-Ming |last13=Qamar |first13=Muhammad Tahir ul |last14=Xia |first14=Wei |last15=Liu |first15=Rui |display-authors=5 |date=2021-11-04 |title=High-quality reference genome sequences of two coconut cultivars provide insights into evolution of monocot chromosomes and differentiation of fiber content and plant height |journal=Genome Biology |volume=22 |issue=1 |page=304 |doi=10.1186/s13059-021-02522-9 |pmc=8567702 |pmid=34736486 |doi-access=free }}</ref> | ||
Another ancestral variety is the '''''niu leka''''' of [[Polynesia]] (the "Compact Dwarfs"). Although it resembles dwarf coconuts (including slow growth), it is genetically distinct and was independently domesticated, likely in [[Tonga]]. Other cultivars of ''niu leka'' may exist in other islands of the Pacific, and some are probably descendants of advanced crosses between Compact Dwarfs and Southeast Asian Dwarf types.<ref name="lebrun2013"/><ref name="Bourdeix"/> | |||
== Distribution and habitat == | |||
Coconuts have a nearly [[cosmopolitan distribution]] due to human cultivation and dispersal. However, their original distribution was in the [[Central Indo-Pacific]], in the regions of [[Maritime Southeast Asia]] and [[Melanesia]].<ref name="Perera"/> | |||
The coconut palm thrives on sandy soils and is highly tolerant of [[salinity]]. It prefers areas with abundant sunlight and regular rainfall of between 1500 mm and 2500 mm per year. It prefers humidity above 60%. If rainfall is less than this, it can survive if its roots can reach the soil water table, but it cannot tolerate waterlogging. It grows from sea level to an altitude of 600 metres in the tropics. It can tolerate a dry season of one month on sandy soils inland, and as much as three months on heavier soils, but the soil must be free-draining. It grows on soils with a pH of 4.5 to 8 (the latter on coral atolls), but prefers a range of 5.5 to 7. Growth is seriously limited by shade. It can resist hurricane-strength winds provided it has developed a good root system.<ref name="sppia">{{cite book |last1=Chan |first1=Edward |first2=Craig R. |last2=Elevitch |date=April 2006 |url=http://www.agroforestry.net/tti/Cocos-coconut.pdf |chapter=Cocos nucifera (coconut): Environmental Preferences and Tolerances |pages=9–10 |archive-url=https://web.archive.org/web/20131020005217/http://agroforestry.net/tti/Cocos-coconut.pdf |archive-date=October 20, 2013 |url-status=live |editor=C. R. Elevitch |title=Species Profiles for Pacific Island Agroforestry |location=Hōlualoa, Hawaii |publisher=Permanent Agriculture Resources}}</ref> | |||
Wild coconuts are restricted to coastal areas in sandy, saline soils. The fruit is adapted for ocean dispersal. Coconuts could not reach inland locations without human intervention to carry seednuts and plant seedlings.<ref name=HCH_2>{{cite journal |last1=Harries |first1=H. C. |year=2012 |title=Germination rate is the significant characteristic determining coconut palm diversity |journal=Annals of Botany |volume =2012 |article-number=pls045 |doi=10.1093/aobpla/pls045 |pmid=23275832 |pmc=3532018 }}</ref> | |||
== Cultivation == | |||
[[File:Efate East - Very old coconut plantations - panoramio.jpg|thumb|A coconut plantation in [[Efate]], [[Vanuatu]]]] | |||
Coconut palms are normally cultivated in hot and wet tropical climates. They need year-round warmth and moisture to grow well and fruit. Coconut palms are hard to establish in dry climates, and cannot grow there without frequent irrigation. In drought conditions, the new leaves do not open well, older leaves may become desiccated, and fruit may be shed.<ref name="sppia"/> | |||
The extent of cultivation in the tropics is threatening a number of habitats, such as [[mangrove]]s; an example of such damage to an ecoregion is in the Petenes mangroves of the [[Yucatán region|Yucatán]].<ref>{{cite web |url=http://www.eoearth.org/article/Petenes_mangroves?topic=49597 |archive-url=https://web.archive.org/web/20121011011735/http://www.eoearth.org/article/Petenes_mangroves?topic=49597 |archive-date=October 11, 2012 |title=Petenes mangroves: types and severity of threats |last1=McGinley |first1=Mark |last2=Hogan |first2=C. Michael |series=The Encyclopedia of Earth |publisher=[[World Wide Fund for Nature]] |date=19 April 2011 |access-date=17 December 2016 }}</ref> Uniquely among trees, coconut trees can be irrigated with sea water.<ref>{{cite book |last=Carr |first=M. K. V. |title=Advances in irrigation agronomy: plantation crops |date=2012-04-05 |publisher=[[Cambridge University Press]] |page=101}}</ref> | |||
=== Pests and diseases === | |||
{{further|List of coconut palm diseases}} | {{further|List of coconut palm diseases}} | ||
[[File:Coconutlethalyellowing.jpg|thumb|upright|A coconut palm dying of [[lethal yellowing]], a [[phytoplasma]] disease]] | |||
Coconuts are susceptible to the [[phytoplasma]] disease, [[lethal yellowing]]. Yellowing diseases affect plantations in Africa, India, Mexico, the Caribbean and the [[Pacific Region]].<ref name="Bourdeix 2016">{{cite web |last=Bourdeix |first=Ronald |url=https://asiatimes.com/article/future-coconut-threatened-lethal-yellowing-disease/ |title=Clarion call for King Coconut |date=9 December 2016 |website=www.atimes.com |access-date=14 December 2016}}</ref> | |||
The coconut palm is damaged by the [[larva]]e of many [[Lepidoptera]] ([[butterfly]] and [[moth]]) species which feed on it, including the [[African armyworm]] (''Spodoptera exempta'') and ''[[Batrachedra]]'' spp.: ''B. arenosella'', ''B. atriloqua'' (feeds exclusively on ''C. nucifera''), ''B. mathesoni'' (feeds exclusively on ''C. nucifera''), and ''B. nuciferae''.<ref>{{cite journal |last1=Yarro |first1=J. G. |last2=Otindo |first2=B. L. |last3=Gatehouse |first3=A. G. |last4=Lubega |first4=M. C. |date=December 1981 |title=Dwarf variety of coconut, ''Cocos nucifera'' (Palmae), a hostplant for the African armyworm, Spodoptera exempta (Wlk.) (Lepidoptera, Noctuidae)|journal=International Journal of Tropical Insect Science |volume=1 |issue=4 |pages=361–362 |doi=10.1017/S1742758400000667 |bibcode=1981IJTIS...1..361Y |s2cid=88007323}}</ref> | |||
The coconut leaf beetle ''[[Brontispa longissima]]'' feeds on young leaves, and damages both [[seedlings]] and mature coconut palms. In 2007, the Philippines imposed a [[quarantine]] in [[Metro Manila]] and 26 provinces to stop the spread of the [[pest (organism)|pest]] and protect the Philippine coconut industry managed by some 3.5 million farmers.<ref>{{cite web |url=http://www.gmanetwork.com/news/story/62343/news/regions/report-26-provinces-quarantined-for-coconut-pest |title=Report: 26 provinces quarantined for coconut pest |publisher=GMA News Online |date=28 September 2007 |access-date=17 December 2016 |url-status=live |archive-url=https://web.archive.org/web/20161220190900/http://www.gmanetwork.com/news/story/62343/news/regions/report-26-provinces-quarantined-for-coconut-pest |archive-date=December 20, 2016 }}</ref> | |||
</ref> | |||
The | The fruit may be damaged by [[eriophyid]] [[coconut mite]]s (''[[Aceria guerreronis]]''). This mite infests coconut plantations, and can cause economic damage up to 60% of coconut production.<ref name="CABI">{{cite web |title=''Aceria guerreronis'' (coconut mite) |url=https://www.cabi.org/isc/datasheet/2596 |access-date=22 November 2025 |work=Invasive Species Compendium |publisher=[[CAB International|CABI]]}}</ref> The immature seeds are infested and damaged by larvae.<ref>{{cite thesis |last=McDonald |first=Sharon Angella |title=Infestation patterns of the coconut mite, ''Eriophyes guerreronis'' (Keifer) (Acari: Eriophyidae), on coconuts and resulting yield loss in eastern Jamaica |date=July 9, 1997 |access-date=2025-11-22 |degree=[[Master of Science]] |publisher=[[Virginia Polytechnic Institute and State University]] |url=http://hdl.handle.net/10919/36916 |hdl=10919/36916}}</ref> Chemical control is possible, but since it needs to be repeated frequently it is impracticable on grounds of cost, environmental harm, and [[pesticide residue]]s in coconut meat and coconut water.<ref name=Lindquist>{{cite book|author1=Lindquist, E.E.|author2=Bruin, J.|author3=Sabelis, M.W.|title=Eriophyoid Mites: Their Biology, Natural Enemies and Control |url=https://books.google.com/books?id=MVmTn4ghwo0C&pg=PA561 |year=1996 |publisher=[[Elsevier]] |isbn=978-0-08-053123-6 |pages=561–569}}</ref> | ||
=== Cultivars === | |||
[[File:Young coconut of Srilanka.jpg|thumb|upright=0.6|[[King coconut]], a shorter cultivar of Sri Lanka]] | |||
Coconut has many commercial and traditional [[cultivar]]s. They can be sorted mainly into tall, dwarf, and hybrid cultivars (hybrids between talls and dwarfs).<ref>{{cite journal |last1=Parthasarathy |first1=V. I. |last2=Geethalakshmi |first2=P.U. |last3=Niral |first3=V. I. |title=Analysis of coconut cultivars and hybrids using isozyme polymorphism |journal=Acta Botanica Croatica |date=15 April 2004 |volume=63 |issue=1 |pages=69-74 |url=https://hrcak.srce.hr/file/5620}}</ref> Varieties are often regional, such as Ceylon Tall, Jamaica Tall, Java Tall, and Malayan Tall.<ref name="Elzabroek"/> | |||
== | Dwarf varieties of Pacific coconuts have been cultivated by [[Austronesian peoples]] since ancient times. These varieties were selected for slower growth, sweeter coconut water, and often brightly colored fruits.<ref name="lebrun2013">{{cite book |last1=Lebrun |first1=P. |last2=Grivet |first2=L. |last3=Baudouin |first3=L. |title=Current Advances in Coconut Biotechnology |publisher=[[Springer Science & Business Media]]|year=2013 |isbn=978-94-015-9283-3 |editor1-last=Oropeza |editor1-first=C. |pages=83–85 |chapter=Use of RFLP markers to study the diversity of the coconut palm |editor2-last=Verdeil |editor2-first=J.K. |editor3-last=Ashburner |editor3-first=G. R. |editor4-last=Cardeña |editor4-first=R. |editor5-last=Santamaria |editor5-first=J. M.}}</ref> Dwarf varieties include Dwarf Green and Dwarf Orange.<ref name="Elzabroek"/> | ||
Varieties have been selected for diverse traits: for example, [[King coconut]] is a Sri Lankan variety with a relatively low sugar content, while [[Macapuno]] has soft jelly-like flesh that fills the whole central cavity; it is used to make sweet desserts.<ref>{{cite journal |last1=Sukendah |last2=Volkaert |first2=Hugo |last3=Sudarsono |date=2009 |title=Isolation and Analysis of DNA Fragment of Genes Related to Kopyor Trait in Coconut Plant |url=https://jurnal.ugm.ac.id/ijbiotech/article/view/7814/6055 |journal=Indonesian Journal of Biotechnology |volume=14 |issue=2 |pages=1169–1178}}</ref><ref name="Florida Gardener">{{cite web |title=Coconut Varieties |url=http://www.floridagardener.com/palms/coconutpalm.htm |url-status=live |archive-url=https://web.archive.org/web/20151020105211/http://www.floridagardener.com/palms/coconutpalm.htm |archive-date=October 20, 2015 |access-date=19 May 2016 |publisher=floridagardener.com }}</ref> | |||
[[Maypan coconut palm|Maypan]] is an [[F1 hybrid]] bred in Jamaica in the 1970s to resist [[lethal yellowing]].<ref>[http://www.avxl82.dsl.pipex.com/CICLY/main.html Centre for Information on Coconut Lethal Yellowing (CICLY)] {{webarchive|url=https://web.archive.org/web/20090605224039/http://www.avxl82.dsl.pipex.com/CICLY/main.html |date=2009-06-05 }}</ref><ref>{{cite journal |last1=Harries |first1=H.C. |last2=Romney |first2=D.H. |year=1974 |title=Maypan: an F1 hybrid coconut variety for commercial production in Jamaica |journal=World Crops |volume=26 |pages=110–111}}</ref> However, Maypan resistance soon began to fail, possibly as early as the 1980s, and certainly by the 2000s.<ref>{{cite journal |last1=Broschat |first1=T. K. |last2=Harrison |first2=N. A. |last3=Donselman |first3=H. |title=Losses to Lethal Yellowing Cast Doubt on Coconut Cultivar Resistance |journal=Palms |date=2002 |volume=46 |issue=4 |pages=185-189 |url=https://palms.org/wp-content/uploads/2016/05/vol46n4p185-189.pdf}}</ref><ref name="Lebrun Baudouin 2007">{{cite journal |last1=Lebrun |first1=P. |last2=Baudouin |first2=L. |last3=Myrie |first3=W. |last4=Berger |first4=A. |last5=Dollet |first5=M. |title=Recent lethal yellowing outbreak: why is the Malayan Yellow Dwarf Coconut no longer resistant in Jamaica? |journal=Tree Genetics & Genomes |volume=4 |issue=1 |date=14 November 2007 |issn=1614-2942 |doi=10.1007/s11295-007-0093-1 |pages=125–131 |url=http://link.springer.com/10.1007/s11295-007-0093-1 |access-date=23 November 2025|url-access=subscription }}</ref> Some other coconut varieties have natural [[crop disease resistance|resistance]] to lethal yellowing with [[allele]]s at [[microsatellite]]s, with ''Vanuatu tall'' and ''Sri-Lanka green dwarf'' as the most resistant cultivars, while ''West African tall'' is especially susceptible.<ref name="Microsatellite">{{cite book |year=2017 |publisher=Springer |first2=S. |first1=M. |last2=Jain |number=17 |last1=Ahuja |title=Biodiversity and Conservation of Woody Plants |doi=10.1007/978-3-319-66426-2 |series=Sustainable Development and Biodiversity |volume=17 |isbn=978-3-319-66425-5 |s2cid=518220 |page=23}}</ref><ref>{{cite book |publisher=Springer |year=2020 |first5=Julianne |first4=Quang |first3=Roland |first2=Mike |first1=Steve |last5=Biddle |last4=Nguyen |last3=Bourdeix |last2=Foale |last1=Adkins |editor-first1=Steve |editor-first2=Mike |editor-first3=Roland |editor-first4=Quang |editor-first5=Julianne |editor-last1=Adkins |editor-last2=Foale |editor-last3=Bourdeix |editor-last4=Nguyen |editor-last5=Biddle |doi=10.1007/978-3-030-44988-9 |title=Coconut Biotechnology: Towards the Sustainability of the 'Tree of Life' |isbn=978-3-030-44987-2 |s2cid=220051653 |pages=128, 132}}</ref><ref>{{cite journal |issue=4 |year=2010 |volume=6 |first6=A. |first5=S. |first4=P. |first3=J.|first2=K. |first1=K. |last6=Sangare |last5=Dery |last4=Lebrun |last3=Konan |last2=Koffi |last1=Konan |title=Microsatellite gene diversity in coconut (''Cocos nucifera'' L.) accessions resistants to lethal yellowing disease |journal=African Journal of Biotechnology |url=https://www.ajol.info/index.php/ajb/article/view/56209}}</ref> | |||
=== Breeding === | |||
Conventional [[plant breeding]] is of limited use with coconut because there is no wild coconut species to supply additional genetic diversity; the generation time is long; there is much [[heterozygosity]]; artificial pollination to conduct crosses produces only a few seeds; and vegetative reproduction (cloning) is unreliable. Coconut breeding objectives can include copra content, production of female flowers, oil content, aroma of coconut meat and water, softness and sweetness of endosperm, drought tolerance, resistance to root wilt, and resistance to eriophorid mites.<ref name="Nair 2016">{{cite book |last1=Nair |first1=Raman V. |url=https://www.researchgate.net/profile/Daniel-Tan-21/publication/305552308_Breeding_Strategies_for_Enhanced_Plant_Tolerance_to_Heat_Stress/links/59d9de68a6fdcc2aad127c82/Breeding-Strategies-for-Enhanced-Plant-Tolerance-to-Heat-Stress.pdf#page=265 |title=Advances in plant breeding strategies: agronomic, abiotic and biotic stress traits |last2=Jerard |first2=B. A. |last3=Thomas |first3=Regi J. |publisher=[[Springer International]] |year=2016 |pages=257-279 |chapter=Coconut breeding in India}}</ref> | |||
===Harvesting=== | Hybrids provide higher precocity and productivity with the number of fruits than other breeds, but they produce fruits with low market acceptance for the water quality of the fruit. Intravarietal crosses in Dwarf coconut (dwarf coconuts bred together) have been tested to provide better water quality for the coconut water market than the hybrids.<ref>{{cite journal |last=Santos |first=Pedro Henrique Araújo Diniz |last2=Venâncio |first2=Thiago Motta |last3=dos Santos |first3=Pedro Henrique Dias |last4=Ramos |first4=Helaine Christine Cancela |last5=Arêdes |first5=Fernanda Abreu Santana |last6=Azevedo |first6=Alinne Oliveira Nunes |last7=Boechat |first7=Marcela Santana Bastos |last8=de Souza Filho |first8=Gonçalo Apolinário |last9=Ramos |first9=Semiramis Rabelo Ramalho |last10=Mirisola |first10=Luiz Angelo |last11=Aragão |first11=Wilson Menezes |last12=Pereira |first12=Messias Gonzaga |display-authors=5 |date=2020 |title=Genotyping-by-sequencing technology reveals directions for coconut (Cocos nucifera L.) breeding strategies for water production |url=https://wikipedialibrary.idm.oclc.org/login?auth=production&url=https://search.ebscohost.com/login.aspx?direct=true&db=a9h&AN=142371411&site=eds-live&scope=site |journal=Euphytica |volume=216 |issue=3 |pages=1–9 |doi=10.1007/s10681-020-02582-1 |access-date=2025-11-22}}</ref> | ||
=== Harvesting === | |||
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| header = Harvesting techniques | |||
| footer = Left: climbing using notches cut into the trunk, [[Philippines]]<br/>Center: using ropes and [[pulley]]s, [[Mexico]]<br/>Right: using a loop of cloth around the ankles, [[Maldives]] | |||
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The two most common harvesting methods are by climbing<ref name="tp2"/> and by using poles.<ref name="cpg"/> | |||
Climbing is the more widespread, but it is more dangerous and requires skilled workers.<ref name="tp2">{{cite web |title=Harvesting and Post-harvest Management |website=Coconut Handbook |date=December 28, 2016 |url=https://coconuthandbook.tetrapak.com/chapter/harvesting-and-post-harvest-management |publisher=[[Tetra Pak]] |access-date=18 April 2021}}</ref> Manually climbing trees is traditional in most countries and requires a posture that exerts pressure on the trunk with the feet. Climbers employed on coconut plantations often develop musculoskeletal disorders and risk injury or death from falling.<ref name="Piggott">{{cite book |last1=Piggott |first1=C.J. |title=Coconut Growing |date=1964 |publisher=[[Oxford University Press]] |page=61}}</ref><ref>{{cite web |title=Tropical Tree and Palm Crops |url=https://www.iloencyclopaedia.org/part-x-96841/agriculture-and-natural-resources-based-industries/tree-bramble-and-vine-crops/item/548-tropical-tree-and-palm-crops |website=Encyclopedia of Occupational Health & Safety |publisher=[[International Labour Organization]] |access-date=24 April 2021}}</ref><ref name="George">{{cite journal |last1=George |first1=Bincy M. |last2=Rao |first2=Muddanna S. |last3=Kumar |first3=Arunachalam |last4=Suvarna |first4=Niveditha |last5=D'Souza |first5=Jessica Sushima |title=Health of coconut tree climbers of rural southern India – medical emergencies, body mass index and occupational marks: A quantitative and survey study |journal=Journal of Clinical and Diagnostic Research |date=2012 |volume=6 |issue=1 |pages=57–60 |url=https://www.jcdr.net/articles/pdf/1829/13%20-%203430.(A).pdf}}</ref> For safety, coconut climbers in the Philippines and [[Guam]] use a [[bolo knife]] tied with a rope to the waist to cut grooves at regular intervals on the coconut trunks. This makes the trunk of the tree more like a ladder, though it reduces the value of timber recovered from the trees and can admit infection.<ref name="fao-timber">{{cite book |title=Coconut wood: Processing and Use |date=1985 |publisher=Food and Agriculture Organization of the United Nations |isbn=92-5-102253-4 |page=27 |url=http://www.fao.org/3/an792e/an792e00.pdf}}</ref><ref name="tp2"/><ref name="Safford">{{cite journal |last=Safford |first=William Edwin |title=The Useful Plants of the Island of Guam |journal=[[Bulletin of the United States National Museum]] |date=8 April 1905 |volume=9 |page=243}}</ref> Other methods to make climbing easier include using a system of pulleys and ropes; using pieces of vine, rope, or cloth tied to both hands or feet; using spikes attached to the feet or legs; or attaching coconut husks to the trunk with ropes.<ref name="COGENT"/> | |||
[[File:“Lambanog”- Philippines Coconut Wine.jpg|thumb|Worker in the [[Philippines]] using a bamboo bridge network to collect sweet coconut sap from cut flower stalks to make [[lambanog|palm liquor]] ]] | |||
The pole method uses a long pole with a cutting device at the end. In the Philippines, the traditional tool is called the ''halabas'' and is made from a long bamboo pole with a sickle-like blade at its tip. Though safer and faster than climbing, it does not allow workers to examine and clean the crown of coconuts for pests and diseases.<ref name="cpg">{{cite web |title=Coconut Production Guide |url=https://businessdiary.com.ph/3062/coconut-production-guide/#Harvest_Management |website=Business Diary PH |date=March 30, 2020 |access-date=18 April 2021}}</ref> | |||
Modern methods use hydraulic elevators mounted on tractors or ladders.<ref name="Loyola"/> Mechanical coconut climbing devices and robots have been developed in India, Sri Lanka, and [[Malaysia]].<ref name="tp">{{cite book |author=Tetra Pak |title=Coconut Handbook |date=2016 |publisher=Tetra Pak International S.A. |isbn=978-91-7773-948-7 |chapter=The Chemistry of Coconut Milk and Cream |chapter-url=https://coconuthandbook.tetrapak.com/chapter/chemistry-coconut-milk-and-cream}}</ref><ref name="Kumar">{{cite news |last1=Kumar |first1=V. Sajeev |title=New mechanical coconut climbing device developed |url=https://www.thehindubusinessline.com/economy/agri-business/New-mechanical-coconut-climbing-device-developed/article20111199.ece |access-date=18 April 2021 |work=The Hindu Business Line |date=20 March 2011}}</ref><ref name="Subramanian">{{cite journal |last1=Subramanian |first1=Parvathi |last2=Sankar |first2=Tamil Selvi |title=Development of a novel coconut-tree-climbing machine for harvesting |journal=Mechanics Based Design of Structures and Machines |date=7 April 2021 |volume=51 |issue=5 |pages=2757–2775 |doi=10.1080/15397734.2021.1907756|s2cid=233530078 }}</ref><ref name="COGENT">{{cite web |title=Climbing the coconut palm |url=http://cogentnetwork.org/oldsite/climbing-the-coconut-palm |website=COGENT |access-date=24 April 2021 |archive-date=December 16, 2021 |archive-url=https://web.archive.org/web/20211216083837/http://cogentnetwork.org/oldsite/climbing-the-coconut-palm }}</ref> The Coconut Maturity Detection Project uses imaging and [[machine learning]] to identify mature coconut bunches ready for harvesting.<ref>{{cite journal |last=Geethalaxmi |last2=Anjali |last3=Ashritha |last4=Sharma |first4=Rajan |last5=Bhat |first5=Shilpa Ganapati |date=2024-06-15 |title=Real Time Sustainable Cultivation of Coconut Tree Crops using ML |url=https://wikipedialibrary.idm.oclc.org/login?auth=production&url=https://search.ebscohost.com/login.aspx?direct=true&db=aci&AN=181715044&site=eds-live&scope=site |journal=Grenze International Journal of Engineering & Technology (GIJET) |volume=10 |issue=2,Part 4 |pages=4666–4670 |access-date=2025-11-22}}</ref> | |||
[[File:Tanjong Jara (4280556051) (2).jpg|thumb|A [[Southern pig-tailed macaque]] trained to harvest coconuts, Malaysia]] | |||
A system of bamboo bridges and ladders directly connecting the tree canopies is used in the Philippines for coconut plantations that harvest coconut sap (not fruits) for [[coconut vinegar]] and [[palm wine]] production.<ref name="Bello">{{cite journal |last1=Bello |first1=Rolando T. |last2=Pantoja |first2=Blanquita R. |last3=Tan |first3=Maria Francesca O. |last4=Banalo |first4=Roxanne A. |last5=Alvarez |first5=Jaonne V. |last6=Rañeses |first6=Florita P. |title=A study on skills for trade and economic diversification (STED) in the non-traditional coconut export sectors of the Philippines |journal=Employment Working Paper |date=2020 |publisher=[[International Labour Organization|International Labor Organization]] |location=Geneva |page=18 |url=https://www.ilo.org/wcmsp5/groups/public/---ed_emp/documents/publication/wcms_742455.pdf}}</ref><ref name="Loyola"/> In other areas, as in [[Papua New Guinea]], coconuts are simply collected when they fall to the ground.<ref name="tp2"/> | |||
=== | Some coconut farmers in [[Thailand]] and Malaysia use [[southern pig-tailed macaque]]s to harvest coconuts.<ref>{{cite journal |last1=Bertrand |first1=M |title=Training without reward: traditional training of pig-tailed macaques as coconut harvesters. |journal=Science |date=27 January 1967 |volume=155 |issue=3761 |pages=484–6 |doi=10.1126/science.155.3761.484 |pmid=4959310 |bibcode=1967Sci...155..484B }}</ref> Thailand has been raising and training them to pick coconuts for around 400 years.<ref name="npr">{{cite web |last=Barclay |first=Eliza |date=October 19, 2015 |title=What's Funny About The Business Of Monkeys Picking Coconuts? |url=https://www.npr.org/sections/thesalt/2015/10/19/448960760/monkeys-pick-coconuts-in-thailand-are-they-abused-or-working-animals |website=[[NPR]]}}</ref><ref name="vm">{{cite news |title=The truth behind the "widespread" practice of monkeys harvesting coconuts in Thailand impacting on consumers & retailers |url=https://vegworldmag.com/the-truth-behind-the-widespread-practice-of-monkeys-harvesting-coconuts-in-thailand-impacting-on-consumers-retailers/ |access-date=18 April 2021 |work=Vegworld Magazine |date=31 July 2020}}</ref><ref name="Pratruangkrai">{{cite news |last1=Pratruangkrai |first1=Petchanet |title=Focus: Thai coconut farmers reject monkey abuse charges, suffer lost income |url=https://english.kyodonews.net/news/2020/07/438857ee536a-focus-thai-coconut-farmers-reject-monkey-abuse-charges-suffer-lost-income.html |access-date=18 April 2021 |work=Kyodo News |date=16 July 2020}}</ref> Training schools for pig-tailed macaques still exist in southern Thailand and in the Malaysian state of [[Kelantan]].<ref>{{cite journal |last=Bertrand |first=Mireille |date=January 27, 1967 |title=Training without Reward: Traditional Training of Pig-tailed Macaques as Coconut Harvesters |journal=[[Science (journal)|Science]] |volume=155 |issue=3761 |pages=484–486 |doi=10.1126/science.155.3761.484}}</ref> [[People for the Ethical Treatment of Animals]] (PETA) objected to the Thailand practice in 2019.<ref name="Loyola">{{cite news |last=Loyola |first=James A. |title=Philippine coconut industry: No monkey business here |url=https://mb.com.ph/2020/07/23/philippine-coconut-industry-no-monkey-business-here/ |access-date=18 April 2021 |work=[[Manila Bulletin]] |date=23 July 2020}}</ref> | ||
{{ | |||
{| class="wikitable" style=" | {| class="wikitable floatright" style="width:12em; text-align:center;" | ||
|+ Coconut production | |+ Coconut (in shell) production <br />{{small|2023, millions of tonnes}}<br/> | ||
|- | |- | ||
|{{IDN}} || | |{{IDN}} ||18.0 | ||
|- | |- | ||
|{{PHL}} || 14.9 | |{{PHL}} ||14.9 | ||
|- | |- | ||
|{{IND}} || | |{{IND}} ||14.2 | ||
|- | |- | ||
|{{BRA}} || 2. | |{{BRA}} ||2.9 | ||
|- | |- | ||
|{{LKA}} || 2. | |{{LKA}} ||2.1 | ||
|- | |- | ||
! '''World''' !! | ! '''World''' !! 64.7 | ||
|- | |- | ||
|colspan=2| | |colspan=2|{{small|Source: [[FAOSTAT]] of the [[United Nations]]}}<ref name="faostat">{{cite web |url=https://www.fao.org/faostat/en/#data/QCL |title=Coconut production in 2023, Crops/Regions/World list/Production Quantity/Year (pick lists) |publisher=UN Food and Agriculture Organization, Corporate Statistical Database |date=2025 |access-date=23 November 2025}}</ref> | ||
|} | |} | ||
=== Production === | |||
{{Main|List of countries by coconut production}} | |||
{{See also|Asian and Pacific Coconut Community}} | |||
In 2023, world production of coconuts in shells was 65 million [[tonne]]s, led by [[Indonesia]], India, and the Philippines, with 73% of the total when combined (table). | |||
In India, four southern states account for most of India's total production: [[Tamil Nadu]], [[Karnataka]], [[Kerala]], and [[Andhra Pradesh]].<ref>{{cite web |url=http://www.coconutboard.gov.in/stat.htm |title=Coconut Cultivation |author1=Coconut Development Board, Government of India |date=n.d. |access-date=December 8, 2015 |archive-url=https://web.archive.org/web/20151210194230/http://www.coconutboard.gov.in/stat.htm |archive-date=December 10, 2015 }}</ref> Though Kerala has the largest number of coconut trees, Tamil Nadu is the most productive per hectare.<ref>{{cite web |url=http://www.coconutboard.nic.in/cps-tn.htm |title=Coconut Cultivation |author=Coconut Development Board, Government of India |date=n.d. |access-date=December 8, 2015 |archive-url=https://web.archive.org/web/20151208153858/http://www.coconutboard.nic.in/cps-tn.htm |archive-date=December 8, 2015 }}</ref> The coconut is the official state tree of Kerala, whose name in the local language [[Malayalam]] means "coconut land".<ref>{{cite web |title=Kerala at a glance |url=https://forest.kerala.gov.in/en/kerala-at-a-glance/ |publisher=[[Kerala Forest Department]] |access-date=21 November 2025}}</ref> | |||
The main coconut-producing area in the Middle East is the [[Dhofar]] region of [[Oman]].<ref>{{cite journal |last1=Perera |first1=Lalith |last2=Baudouin |first2=Luc |last3=Bourdeix |first3=Roland |last4=Fadhil |first4=Anwar Bait |last5=Hountondji |first5=Fabien C. C. |last6=Al-Shanfri |first6=Abdulaziz |title=Coconut Palms on the Edge of the Desert: Genetic Diversity of Cocos nucifera L. in Oman |journal=Cord |date=2011 |volume=27 |issue=1 |pages=9-19 |url=https://agritrop.cirad.fr/560027/1/120-Article%20Text-198-1-10-20200313.pdf}}</ref> The ancient coconut groves of Dhofar were mentioned by the medieval Moroccan traveller [[Ibn Battuta]] in his book ''[[The Rihla]]''.<ref>{{cite web |url=http://www.fordham.edu/halsall/source/1354-ibnbattuta.html |title=Medieval Sourcebook: Ibn Battuta: Travels in Asia and Africa 1325–1354 |editor-last=Halsall |editor-first=Paul |publisher=[[Fordham University]] Center for Medieval Studies |date=February 21, 2001 |access-date=April 14, 2011 |archive-url=https://web.archive.org/web/20110513172555/http://www.fordham.edu/halsall/source/1354-ibnbattuta.html |archive-date=May 13, 2011 }}</ref> | |||
[[Sri Lanka]] is the fourth-largest producer, and is the second-largest producer of coconut oil and copra, accounting for 15% of global production. The production of coconuts is the main source of [[Economy of Sri Lanka|Sri Lanka economy]], with 12% of cultivated land and 409,244 hectares used for coconut growing (2017). Sri Lanka established its Coconut Development Authority and Coconut Cultivation Board and Coconut Research Institute in the early [[British Ceylon]] period.<ref name="sri">{{cite web |title=Coconut Industry in Sri Lanka |publisher=Sri Lanka Export Development Board |url=https://www.srilankabusiness.com/coconut/about/#:~:text=Coconut%20accounts%20for%20approximately%2012,3600%20million%20nuts%20per%20year. |access-date=January 3, 2021 |website=Sri Lanka Business}}</ref> | |||
[[Sri Lanka]] is the | |||
==== | == Health concerns == | ||
=== Cardiovascular disease === | |||
{{further|Coconut oil#Health concerns|Saturated fat#Cardiovascular disease}} | |||
==== | As a rich source of lauric acid and other saturated fats, coconut oil is a dietary [[risk factor]] for [[cardiovascular disease]]s by elevating blood levels of [[low-density lipoprotein|LDL cholesterol]].<ref name="aha">{{cite journal |author=Neelakantan, N.; Seah, J.Y.; van Dam, R.M. |title=The Effect of Coconut Oil Consumption on Cardiovascular Risk Factors: A Systematic Review and Meta-Analysis of Clinical Trials |journal=Circulation |volume=141 |issue=10 |pages=803–814 |date=March 2020 |pmid=31928080 |doi=10.1161/CIRCULATIONAHA.119.043052 |url=https://www.ahajournals.org/doi/10.1161/CIRCULATIONAHA.119.043052|quote=Coconut oil should not be viewed as healthy oil for cardiovascular disease risk reduction, and limiting coconut oil consumption because of its high saturated fat content is warranted.|url-access=subscription }}</ref><ref name="spiazzi">{{cite journal |last1=Spiazzi |first1=B.F. |last2=Duarte |first2=A.C. |last3=Zingano |first3=C.P. |display-authors=etal |title=Coconut oil: an overview of cardiometabolic effects and the public health burden of misinformation |journal=Archives of Endocrinology and Metabolism |volume=67 |issue=6 |article-number=e000641 |date=June 2023 |pmid=37364144 |pmc=10660992 |doi=10.20945/2359-3997000000641|quote=The use of coconut oil as a "healthy" component of the western diet is based on the major spread of misconceptions regarding it. The combination of the established knowledge on the negative effects of saturated fats on cardiovascular health and the lack of evidence from clinical trials showing a benefit from coconut oil intake in cardiovascular and metabolic disease advise against the consumption of this oil as a preferential source of dietary fat.}}</ref> Many national clinical associations recommend limiting consumption of coconut oil products, replacing them instead with foods containing [[unsaturated fat]]s.<ref name=aha/><ref name=spiazzi/><ref name="aus">{{cite web |title=Sorting fact from fiction – seed oils and coconut oil |url=https://www.heartfoundation.org.au/blog/seed-oils-coconut-oil |publisher=[[National Heart Foundation of Australia]] |access-date=23 November 2025 |date=23 May 2025}}</ref> | ||
== Allergens == | === Allergens === | ||
Proteins from coconut [[Food allergy|in food may cause allergic reactions]], including [[anaphylaxis]], in some people.<ref name="MichavilaAmat2015">{{cite journal |last1=Michavila Gomez |first1=A. |last2=Amat Bou |first2=M. |last3=Gonzalez Cortés |first3=M.V. |last4=Segura Navas |first4=L. |last5=Moreno Palanques |first5=M.A. |last6=Bartolomé |first6=B. |title=Coconut anaphylaxis: Case report and review |journal=Allergologia et Immunopathologia |volume=43 |issue=2 |date=2015 |doi=10.1016/j.aller.2013.09.004 |pages=219–220 |url=https://linkinghub.elsevier.com/retrieve/pii/S0301054613002449 |access-date=23 November 2025|doi-access=free }}</ref> In the United States, the [[Food and Drug Administration]] has declared that coconut must be disclosed as an ingredient on food package labels as a "tree nut" with potential [[allergen]]icity.<ref>{{cite web |date=January 2013 |title=Guidance for Industry: A Food Labeling Guide (6. Ingredient Lists); Major Food Allergens (food source names and examples) |url=https://www.fda.gov/food/guidanceregulation/guidancedocumentsregulatoryinformation/labelingnutrition/ucm064880.htm |archive-url=https://web.archive.org/web/20160529191639/https://www.fda.gov/Food/GuidanceRegulation/GuidanceDocumentsRegulatoryInformation/LabelingNutrition/ucm064880.htm |archive-date=May 29, 2016 |access-date=29 May 2016 |work=Food Labeling and Nutrition |publisher=[[Food and Drug Administration]] }}</ref> | |||
[[ | |||
[[Cocamidopropyl betaine]] (CAPB) is a [[surfactant]] manufactured from coconut oil and used in personal hygiene products and cosmetics, such as [[shampoo]]s, liquid [[soap]]s, cleansers and [[antiseptic]]s. CAPB may cause mild [[skin irritation]],<ref name="JacobAmini2008">{{cite journal |last1=Jacob |first1=S. E. |last2=Amini |first2=S. |year=2008 |title=Cocamidopropyl betaine |journal=Dermatitis |type=Review |volume=19 |issue=3 |pages=157–160 |doi=10.2310/6620.2008.06043 |pmid=18627690}}</ref> but allergic reactions to CAPB are rare;<ref name="SchnuchLessmann2011">{{cite journal |last1=Schnuch |first1=A. |last2=Lessmann |first2=H. |last3=Geier |first3=J. |last4=Uter |first4=W. |year=2011 |title=Is cocamidopropyl betaine a contact allergen? Analysis of network data and short review of the literature |journal=Contact Dermatitis |type=Review |volume=64 |issue=4 |pages=203–211 |doi=10.1111/j.1600-0536.2010.01863.x |pmid=21392028 |s2cid=205814762}}</ref> they are probably related to impurities rendered during the manufacturing process (which include [[amidoamine]] and [[dimethylaminopropylamine]]) rather than CAPB itself.<ref name="JacobAmini2008"/> | |||
[[Cocamidopropyl betaine]] (CAPB) is a [[surfactant]] manufactured from coconut oil | |||
== Uses == | == Uses == | ||
{{main|Uses of the coconut palm}} | |||
It is one of the most useful trees in the world.<ref name="duke" /> | The coconut palm is grown throughout the [[tropics]] for decoration, as well as for its culinary and nonculinary uses; virtually [[uses of the coconut palm|every part of the coconut palm is used by humans]] in some manner and has significant economic value. The coconut's versatility is noted in some of its names: in [[Sanskrit]], it is ''kalpa vriksha'' ("the tree of the necessities of life"), while in [[Malay language|Malay]], it is ''pokok seribu guna'' ("the tree of a thousand uses"), and in the Philippines, it is called the "[[tree of life]]".<ref>{{cite web |last=Margolis |first=Jason |date=December 13, 2006 |url=http://www.theworld.org/?q=node/7280 |title=Coconut fuel |archive-url=https://web.archive.org/web/20110831035256/http://www.theworld.org/?q=node%2F7280 |archive-date=August 31, 2011 |website=PRI's The World |access-date=April 10, 2011}}</ref> It is one of the most useful trees in the world.<ref name="duke"/> | ||
=== | === Nutrition === | ||
{{Nutritional value | {{Nutritional value | ||
| name=Coconut meat, raw | |name=Coconut meat, raw | ||
|image=Coconuts - single and cracked open.jpg | |image=Coconuts - single and cracked open.jpg | ||
| water=47 g | |water=47 g | ||
| kJ=1480 | |kJ=1480 | ||
| protein=3.33 g | |protein=3.33 g | ||
| fat=33.49 g | |fat=33.49 g | ||
| satfat=29.698 g | |satfat=29.698 g | ||
| monofat = 1.425 g | |monofat=1.425 g | ||
| polyfat = 0.366 g | |polyfat=0.366 g | ||
| carbs=15.23 g | |carbs=15.23 g | ||
| sugars=6.23 g | |sugars=6.23 g | ||
| fiber=9.0 g | |fiber=9.0 g | ||
| calcium_mg=14 | |calcium_mg=14 | ||
| copper_mg=0.435 | |copper_mg=0.435 | ||
| iron_mg=2.43 | |iron_mg=2.43 | ||
| magnesium_mg=32 | |magnesium_mg=32 | ||
| phosphorus_mg=113 | |phosphorus_mg=113 | ||
| potassium_mg=356 | |potassium_mg=356 | ||
| sodium_mg=20 | |sodium_mg=20 | ||
| zinc_mg=1.10 | |zinc_mg=1.10 | ||
| manganese_mg=1.500 | |manganese_mg=1.500 | ||
| selenium_ug=10.1 | |selenium_ug=10.1 | ||
| vitC_mg=3.3 | |vitC_mg=3.3 | ||
| thiamin_mg=0.066 | |thiamin_mg=0.066 | ||
| riboflavin_mg=0.020 | |riboflavin_mg=0.020 | ||
| niacin_mg=0.540 | |niacin_mg=0.540 | ||
| pantothenic_mg=0.300 | |pantothenic_mg=0.300 | ||
| vitB6_mg=0.054 | |vitB6_mg=0.054 | ||
| folate_ug=26 | |folate_ug=26 | ||
| vitE_mg=0.24 | |vitE_mg=0.24 | ||
| vitK_ug=0.2 | |vitK_ug=0.2 | ||
<!-- amino acids --> | <!-- amino acids --> | ||
| tryptophan=0.039 g | |tryptophan=0.039 g | ||
| threonine=0.121 g | |threonine=0.121 g | ||
| isoleucine=0.131 g | |isoleucine=0.131 g | ||
| leucine=0.247 g | |leucine=0.247 g | ||
| lysine=0.147 g | |lysine=0.147 g | ||
| methionine=0.062 g | |methionine=0.062 g | ||
| cystine=0.066 g | |cystine=0.066 g | ||
| phenylalanine=0.169 g | |phenylalanine=0.169 g | ||
| tyrosine=0.103 g | |tyrosine=0.103 g | ||
| valine=0.202 g | |valine=0.202 g | ||
| arginine=0.546 g | |arginine=0.546 g | ||
| histidine=0.077 g | |histidine=0.077 g | ||
| alanine=0.170 g | |alanine=0.170 g | ||
| aspartic acid=0.325 g | |aspartic acid=0.325 g | ||
| glutamic acid=0.761 g | |glutamic acid=0.761 g | ||
| glycine=0.158 g | |glycine=0.158 g | ||
| proline=0.138 g | |proline=0.138 g | ||
| serine=0.172 g | |serine=0.172 g | ||
|note=[https://fdc.nal.usda.gov/food-details/170169/nutrients Link to USDA FoodData Central entry] | |||
| note=[https://fdc.nal.usda.gov | |||
}} | }} | ||
Raw coconut meat is 47% water, 33% [[fat]], 15% [[carbohydrate]]s, and 3% [[protein (nutrient)|protein]] (table). In a reference amount of {{convert|100|g}}, raw coconut flesh supplies 350 [[calorie]]s of [[food energy]], and is a rich source (20% or more of the [[Daily Value]], DV) of [[manganese in biology|manganese]] (65% DV) and [[copper in biology|copper]] (48% DV), with various other [[mineral (nutrient)|dietary minerals]] in moderate amounts (10-18% DV; table). It is a poor source of [[vitamin]]s. Raw coconut meat has a high content of [[Fatty_acid#Saturated_fatty_acids|saturated fatty acids]] (30% of total fats), with [[lauric acid]] as the main saturated fat (15% of total; USDA source in table). | |||
==== | === Culinary === | ||
{{further|List of coconut dishes}} | |||
{{Cookbook}} | |||
<!--Please do not extend this section: it is a summary of [[Uses of the coconut palm]]--> | |||
The [[List of coconut dishes|many culinary uses of coconuts]] are largely based on the edible white, fleshy part of the seed (the [[endosperm]]), known as "coconut meat".<ref name="Roehl 1996 p. 115"/> The meat of immature coconuts can be eaten as it is or cooked in pastries. Mature coconut meat is tough and is processed before consumption, being made into products like [[coconut milk]],<ref name="Roehl 1996 p. 115"/><ref name="polistico2">{{cite book |author=Edgie Polistico |url=https://books.google.com/books?id=-h9MswEACAAJ |title=Philippine Food, Cooking, & Dining Dictionary |publisher=Anvil Publishing, Incorporated |year=2017 |isbn=978-971-27-3170-9}}</ref><ref>{{cite book |title=Coconut: Pacific food leaflet No. 4 |date=2006 |publisher=Secretariat of the Pacific Community |isbn=978-982-00-0164-0 |url=https://cms.ctahr.hawaii.edu/Portals/159/Images/Resources/IslandStapleFoods/4%20COCONUT%20web.pdf}}</ref><ref name="El Bassam">{{cite book |last1=El Bassam |first1=N. |title=Handbook of Bioenergy Crops: A Complete Reference to Species, Development, and Applications |date=2010 |publisher=Earthscan |isbn=978-1-84977-478-9 |page=159}}</ref><ref>{{cite book |last1=Chu |first1=Wai Hon |last2=Lovatt |first2=Connie |title=The Dumpling: A Seasonal Guide |date=2009 |publisher=Harper Collins |isbn=978-0-06-195940-0 |page=8}}</ref> "coconut chips"<ref name="El Bassam"/> or grated and dehydrated as "desiccated coconut".<ref>{{cite web |title=ICC Quality Standard: Desiccated Coconut |url=https://coconutcommunity.org/viewpdf/apcc_quality_standards_for_coconut_products/4 |website=International Coconut Community |publisher=United Nations Economic and Social Commission for Asia and the Pacific (UN-ESCAP) |access-date=7 June 2021 |archive-date=December 16, 2021 |archive-url=https://web.archive.org/web/20211216085337/https://coconutcommunity.org/viewpdf/apcc_quality_standards_for_coconut_products/4 }}</ref><ref name="Ghosh">{{cite book |last1=Ghosh |first1=D.K. |title=Value Addition of Horticultural Crops: Recent Trends and Future Directions |date=2015 |publisher=Springer |chapter=Postharvest, Product Diversification and Value Addition in Coconut |editor1-first=Amit Baran |editor1-last=Sharangi |editor2-first=Suchand |editor2-last=Datta |isbn=978-81-322-2262-0}}</ref><ref>{{cite web |title=Exporting desiccated coconuts to Europe |url=https://www.cbi.eu/market-information/processed-fruit-vegetables-edible-nuts/desiccated-coconuts/europe |website=CBI |publisher=Centrum tot Bevordering van de Import uit ontwikkelingslanden (CBI), Netherlands Ministry of Foreign Affairs |access-date=7 June 2021}}</ref> | |||
[[Coconut milk]], used for cooking [[List of dishes made using coconut milk|many dishes]], is pressed from coconut meat. It can be diluted to create [[coconut milk beverage]]s such as [[milk substitute]]s.<ref name="NIIR">{{cite book |author=NIIR Board of Consultants and Engineers |url=https://books.google.com/books?id=UvXKAgAAQBAJ&pg=PA274 |title=The Complete Book on Coconut & Coconut Products (Cultivation and Processing) |date=2006 |publisher=Asia Pacific Business Press |isbn=978-81-7833-007-5 |page=274}}</ref><ref name="tp"/> [[Coconut milk powder]], a protein-rich powder, can be processed from coconut milk.<ref>{{cite journal |journal=Applied Biochemistry and Biotechnology |year=2012 |volume=167 |issue=5 |pages=1290–302 |doi=10.1007/s12010-012-9632-9 |title=Production of coconut protein powder from coconut wet processing waste and its characterization |last1=Naik |first1=A. |last2=Raghavendra |first2=S. N. |last3=Raghavarao |first3=K. S. |pmid=22434355 |s2cid=11890199}}</ref> Coconut milk and [[coconut cream]] extracted from grated coconut can be added to desserts and savory dishes, or used in [[curries]] and stews.<ref name="grimwood">[[#Grimwood |Grimwood]], pp. 183–187.</ref><ref name="pca-process">{{cite book |author=Philippine Coconut Authority |title=Coconut Processing Technologies: Coconut Milk |publisher =Department of Agriculture, Republic of the Philippines |series=FPDD Guide No. 2 – Series of 2014 |year=2014 |url=http://www.pca.da.gov.ph/pdf/techno/coconut_milk.pdf |access-date=May 24, 2019 |archive-date=November 6, 2020 |archive-url=https://web.archive.org/web/20201106130020/http://www.pca.da.gov.ph/pdf/techno/coconut_milk.pdf}}</ref> Products made from thickened coconut milk with sugar and eggs, like [[coconut jam]] and [[coconut custard]], are widespread in Southeast Asia.<ref name="Duruz">{{cite book |last1=Duruz |first1=Jean |last2=Khoo |first2=Gaik Cheng |title=Eating Together: Food, Space, and Identity in Malaysia and Singapore |date=2014 |publisher=Rowman & Littlefield |isbn=978-1-4422-2741-5 |page=45 |url=https://books.google.com/books?id=JojpBQAAQBAJ&q=kaya%20coconut&pg=PA45}}</ref><ref name="Alford">{{cite book |last1=Alford |first1=Jeffrey |last2=Duguid |first2=Naomi |title=Hot Sour Salty Sweet: A Culinary Journey Through Southeast Asia |date=2000 |publisher=Artisan Books |isbn=978-1-57965-564-8 |page=302 |url=https://books.google.com/books?id=cUP30wR8-kUC&pg=PA302}}</ref><ref name="thampan 1981 p199">{{cite book |last1=Thampan |first1=Palakasseril Kumaran |title=Handbook on Coconut Palm |date=1981 |publisher=Oxford & IBH |page=199}}</ref> Coconut oil is used for [[frying]] and cooking.<ref name="grimwood"/><ref name="kurian">{{cite book |last1=Kurian |first1=Alice |last2=Peter |first2=K.V. |title=Commercial Crops Technology |date=2007 |publisher=New India Publishing |isbn=978-81-89422-52-3 |pages=202–203 |url=https://books.google.com/books?id=2VFYqwA-Mn4C&pg=PA202}}</ref> | |||
Coconut | [[Coconut water]] can be drunk fresh or used in cooking.<ref name=endosperm>{{cite book |url=https://books.google.com/books?id=cjHCoMQNkcgC&q=coconut+endosperm+encyclopedia&pg=PA112 |title=Cocos in The Encyclopedia of Fruit and Nuts |access-date=11 May 2015 |year=2008 |last1=Janick |first1=J. |last2=Paull |first2=R. E. |pages=109–113 |isbn=978-0-85199-638-7}}</ref><ref>{{cite web |title=Ginataang Manok (Chicken Stewed in Coconut Milk) Filipino Recipe! |url=http://www.savvynana.com/recipe/ginataang-manok-chicken-stewed-coconut-milk-filipino-recipe/ |website=Savvy Nana's |access-date=20 April 2019 |archive-date=December 16, 2021 |archive-url=https://web.archive.org/web/20211216100142/http://www.savvynana.com/recipe/ginataang-manok-chicken-stewed-coconut-milk-filipino-recipe/ |url-status=usurped }}</ref> It can be fermented to produce a jelly-like dessert known as ''[[nata de coco]]''.<ref name="tietze">{{cite book |last1=Tietze |first1=Harald |last2=Echano |first2=Arthur |title=Coconut: Rediscovered as Medicinal Food |date=2006 |publisher=Harald Tietze Publishing |isbn=978-1-876173-57-9 |page=37 |url=https://books.google.com/books?id=0NAA8_fDI9wC&pg=PA37}}</ref> Coconut vinegar, made from fermented coconut water or sap, is used extensively in Southeast Asian and [[Goan cuisine]].<ref name="polistico2"/> | ||
==== | Coconut sap, fresh or fermented, is drunk as toddy or ''[[tubâ]]'' in the Philippines. When left to ferment on its own, it becomes palm wine. Palm wine is distilled to produce ''[[arrack]]''.<ref name="lambanog">{{cite web |url=http://www1.american.edu/ted/lambanog.htm |title=Lambanog: A Philippine Drink |author=Porter, Jolene V. |year=2005 |publisher=American University |location=Washington, D.C. |access-date=April 10, 2011 |archive-url=https://web.archive.org/web/20110222210517/http://www1.american.edu/ted/lambanog.htm |archive-date=February 22, 2011}}</ref> The sap can be reduced by boiling to create a sweet syrup, or reduced further to yield [[coconut sugar]]. A young, well-maintained tree can produce around {{convert |300 |L |USgal |abbr=off}} of toddy per year, while a 40-year-old tree may yield around {{convert |400 |L |USgal |abbr=on}}.<ref>[[#Grimwood |Grimwood]], p. 20.</ref> | ||
==== | ==== Oil ==== | ||
{{main|Coconut oil}} | |||
{| class="wikitable floatright" style="width:13em; text-align:center;" | |||
|+ Coconut oil production <br/>{{small|2022, tonnes}}<br/> | |||
|- | |||
|{{PHL}} ||1,358,900 | |||
= | |- | ||
{{ | |{{IDN}} ||666,000 | ||
|- | |||
|{{IND}} ||360,000 | |||
|- | |||
|{{VIE}} ||182,000 | |||
|- | |||
|{{MEX}} ||132,300 | |||
|- | |||
|{{LKA}} ||84,015 | |||
|- | |||
! '''World''' !! 3,162,279 | |||
|- | |||
|colspan=2|{{small|Source: [[FAOSTAT]]}}<!--ref can't be named "faostat", that clashes with the other faostat ref above--><ref name="faostat OIL">{{cite web |url=https://www.fao.org/faostat/en/#data/QCL |title=Coconut oil production in 2022, Crops/Regions/World list/Production Quantity/Year (pick lists) |publisher=UN Food and Agriculture Organization, Corporate Statistical Database |date=2025 |access-date=23 November 2025}}</ref> | |||
|} | |||
In 2022, world production of processed [[coconut oil]] was 3.2 million tonnes, led by the Philippines with 43% of the total, and Indonesia and India as secondary producers (table). | |||
Coconut oil is used in cooking, especially for frying. It can be used in liquid form like other [[vegetable oil]]s, or in solid form like [[butter]] or [[lard]].<ref>{{cite journal |doi=10.1161/CIR.0000000000000510 |pmid=28620111 |title=Dietary Fats and Cardiovascular Disease: A Presidential Advisory from the American Heart Association |journal=Circulation |volume=136 |issue=3 |pages=e1–e23 |year=2017 |last1=Sacks |first1=Frank M. |last2=Lichtenstein |first2=Alice H. |last3=Wu |first3=Jason H.Y. |last4=Appel |first4=Lawrence J. |last5=Creager |first5=Mark A. |last6=Kris-Etherton |first6=Penny M. |last7=Miller |first7=Michael |last8=Rimm |first8=Eric B. |last9=Rudel |first9=Lawrence L. |last10=Robinson |first10=Jennifer G. |last11=Stone |first11=Neil J. |last12=Van Horn |first12=Linda V. |display-authors=5 |s2cid=367602 |doi-access=free}}</ref><ref name="Neela">{{cite journal |last1=Neelakantan |first1=Nithya |last2=Seah |first2=Jowy Yi Hoong |last3=van Dam |first3=Rob M. |title=The effect of coconut oil consumption on cardiovascular risk factors (Systematic review) |journal=Circulation |volume=141 |issue=10 |date=2020 |doi=10.1161/circulationaha.119.043052 |pages=803–814 |pmid=31928080 |s2cid=210195904 |doi-access=free }}</ref><ref name="Eyres2016">{{cite journal |last1=Eyres |first1=L |last2=Eyres |first2=MF |last3=Chisholm |first3=A |last4=Brown |first4=RC |title=Coconut oil consumption and cardiovascular risk factors in humans |journal=Nutrition Reviews |date=April 2016 |volume=74 |issue=4 |pages=267–280 |doi=10.1093/nutrit/nuw002 |pmid=26946252 |pmc=4892314}}</ref> [[Coconut butter]] is a solidified coconut oil, but the name is also applied to [[creamed coconut]], a specialty product made of coconut milk solids or [[purée]]d coconut meat and oil.<ref name="Roehl 1996 p. 115">{{cite book |last=Roehl |first=E. |title=Whole Food Facts: The Complete Reference Guide |publisher=Inner Traditions/Bear |year=1996 |isbn=978-0-89281-635-4 |url=https://books.google.com/books?id=H8lwmA52ltIC&pg=PA115 |page=115 }}</ref> | |||
==== | === Non-food uses === | ||
[[ | <!--Please do not extend this section: it is a summary of [[Uses of the coconut palm]]--> | ||
Among the many non-food uses of coconut palms, the husk and shells can be used for fuel or made into [[charcoal]].<ref>{{cite web |title=Coconut Shell Lump Charcoal |url=http://www.supremecarbon.com/coco.htm |publisher=Supreme Carbon Indonesia |url-status=live |archive-url=https://web.archive.org/web/20121229141916/http://www.supremecarbon.com/coco.htm |archive-date=December 29, 2012}}</ref> The husks can serve as [[Personal flotation device |flotation devices]] or as an [[abrasive]].<ref name="Skyhorse Publishing">{{cite book |title=The Complete Guide to Edible Wild Plants |publisher=Skyhorse Publishing; United States Department of the Army |year=2009 |isbn=978-1-60239-692-0 |location=New York |page=44 |oclc=277203364}}</ref> The shell, freed from the husk, and heated on warm ashes, exudes an oily material that is used to soothe dental pains in [[traditional medicine]] of Cambodia.<ref name="dyphon">{{cite book |last1=Pauline Dy Phon |author1-link=Pauline Dy Phon |url=https://books.google.com/books?id=InD2RAAACAAJ |title=Plants Utilised In Cambodia |date=2000 |publisher=Imprimerie Olympic |location=Phnon Penh |pages=165–166}}</ref> [[Coir]] fiber from husks is used in [[rope]]s, [[mat]]s, [[brush]]es, and sacks, as [[caulking]] for boats, and as stuffing for [[mattress]]es.<ref>[[#Grimwood |Grimwood]], p. 22.</ref> It is used in [[horticulture]] in potting compost, especially in orchid mix, and to make brooms in Cambodia.<ref name="dyphon" /> [[Coconut cup]]s were frequently carved with scenes in [[relief]] and mounted with precious metals.<ref>{{cite web |title=Hans van Amsterdam: Coconut Cup with Cover (17.190.622ab) – Heilbrunn Timeline of Art History – The Metropolitan Museum of Art |url=http://www.metmuseum.org/toah/works-of-art/17.190.622ab |url-status=live |archive-url=https://web.archive.org/web/20131015101625/http://www.metmuseum.org/toah/works-of-art/17.190.622ab |archive-date=October 15, 2013 |work=metmuseum.org}}</ref> The leaves provide material for [[basket]]s and for roofing [[thatch]]; they can be woven into mats, cooking [[skewer]]s, and kindling [[arrow]]s. Leaves are woven into small pouches that are filled with rice and cooked to make ''[[pusô]]'' and ''[[ketupat]]''.<ref>[[#Grimwood |Grimwood]], p. 19.</ref> | |||
==== | Hawaiians hollowed out coconut trunks to form drums, containers, or small canoes. The "branches" (leaf [[petiole (botany) |petiole]]s) are strong and flexible enough to make a [[switch (rod) |switch]]. The use of coconut branches in corporal punishment was revived in the Gilbertese community on Choiseul in the [[Solomon Islands]] in 2005.<ref>Herming, George. (March 6, 2006). [http://www.corpun.com/sbj00603.htm Wagina whips offenders] {{webarchive |url=https://web.archive.org/web/20061027160604/http://www.corpun.com/sbj00603.htm |date=October 27, 2006 }}. ''Solomon Star''.</ref> The roots are used to make [[dye]], a [[mouthwash]], and a folk medicine for [[diarrhea]] and [[dysentery]].<ref name="Grimwood18">[[#Grimwood|Grimwood]], p. 18.</ref> A frayed piece of root can be used as a [[toothbrush]]. In Cambodia, the roots are used in traditional medicine.<ref name="dyphon" /> Leftover fiber from coconut oil and coconut milk production, coconut meal, is used as livestock feed. The dried [[calyx (botany) |calyx]] is used as fuel in wood-fired [[stove]]s. Coconut water is traditionally used as a growth supplement in [[plant tissue culture]] and [[micropropagation]].<ref>{{cite journal |last1=Yong |first1=J. W. |last2=Ge |first2=L. |last3=Ng |first3=Y. F. |last4=Tan |first4=S. N. |year=2009 |title=The chemical composition and biological properties of coconut (''Cocos nucifera'' L.) water |journal=Molecules |volume=14 |issue=12 |pages=5144–5164 |doi=10.3390/molecules14125144 |pmid=20032881 |pmc=6255029 |doi-access=free }}</ref> | ||
== In culture == | |||
[[File:A worship at the shrine of the Holy Infant Jesus of Prague, Davao City, Philippines.jpg|thumb|''Palaspas'', woven palm fronds during [[Palm Sunday]] celebrations in the Philippines|alt=Group of people with palm fronds]] | [[File:A worship at the shrine of the Holy Infant Jesus of Prague, Davao City, Philippines.jpg|thumb|''Palaspas'', woven palm fronds during [[Palm Sunday]] celebrations in the Philippines|alt=Group of people with palm fronds]] | ||
[[File:Muñeco de coco.jpg|thumb|upright|A "coconut monkey" from [[Mexico]], a common souvenir carved from coconut shells|alt=Coconut carved to look like a monkey]] | [[File:Muñeco de coco.jpg|thumb|upright|A "coconut monkey" from [[Mexico]], a common souvenir carved from coconut shells|alt=Coconut carved to look like a monkey]] | ||
[[File:Offerings on the sand, Kuta Beach, Bali.JPG|thumb|upright|A ''canang'', an offering of flowers, rice, and [[incense]] in woven coconut leaves from [[Bali]], [[Indonesia]]]] | |||
A coconut ({{langx|sa|narikela}}) is used in [[Hindu]] [[rituals]].<ref>{{cite web |title=Coconut – Fruit Of Lustre In Indian Culture|url=http://www.esamskriti.com/essay-chapters/Coconut-~-Fruit-of-Lustre-in-Indian-Culture-1.aspx |date=September 2011 |last1=Patil |first1=Vimla|website=eSamskriti |url-status=live |archive-url=https://web.archive.org/web/20160514010555/http://www.esamskriti.com/essay-chapters/Coconut-~-Fruit-of-Lustre-in-Indian-Culture-1.aspx |archive-date=May 14, 2016 |access-date=May 18, 2016}}</ref> Often it is decorated with bright metal foils. It is offered during worship to a Hindu god or goddess. [[Narali Purnima]] is celebrated on a [[full moon]] day which usually signifies the end of [[monsoon]] season in India. The word ''Narali'' is derived from [[Marathi language|Marathi]] ''naral'', "coconut". Fishermen give an offering of coconut to the sea to celebrate the beginning of a new fishing season,<ref>{{cite web |url=http://www.maharashtratourism.net/festivals/narali-poornima.html |title=Narali Purnima |publisher=Maharashtra Tourism |date=2016 |access-date=August 3, 2016 |url-status=live |archive-url=https://web.archive.org/web/20160715223636/http://www.maharashtratourism.net/festivals/narali-poornima.html |archive-date=July 15, 2016 }}</ref> in the hope of bountiful catches. Hindus often initiate any new activity by breaking a coconut to ensure the blessings of the gods. The Hindu goddess of well-being and wealth, [[Lakshmi]], is often shown holding a coconut.<ref>Dallapiccola, Anna. ''[[Dictionary of Hindu Lore and Legend]]''. {{ISBN|0-500-51088-1}}.</ref> They are used in Hindu weddings as a symbol of prosperity.<ref>{{cite web |date=November 9, 2009 |title=Food Rituals in Hindu Weddings |url=https://www.smithsonianmag.com/arts-culture/food-rituals-in-hindu-weddings-72454431/ |last=Bramen |first=Lisa |website=[[Smithsonian (magazine)|Smithsonian Magazine]] |access-date=May 25, 2020}}</ref> The flowers are used sometimes in wedding ceremonies in Cambodia.<ref name="dyphon"/> The coconut has cultural and religious significance for [[Austronesian peoples]], appearing in their mythologies, songs, and oral traditions.<ref name="Nayar">{{cite book |last=Nayar |first=N. Madhavan |title=The Coconut: Phylogeny, Origins, and Spread |date=2017 |publisher=[[Academic Press]] |isbn=978-0-12-809778-6 |pages=10–21}}</ref><ref name="Lew">{{cite journal |last=Lew |first=Christopher |title=Tracing the origin of the coconut (''Cocos nucifera'' L.) |url=https://prizedwriting.ucdavis.edu/sites/prizedwriting.ucdavis.edu/files/sitewide/Tracing%20the%20origin%20of%20the%20coconut%20%28Cocos%20nucifera%20L.%29.pdf |journal=Prized Writing 2018–2019 |publisher=[[University of California, Davis]] |pages=143–157 |archive-url=https://web.archive.org/web/20210623220429/https://prizedwriting.ucdavis.edu/sites/prizedwriting.ucdavis.edu/files/sitewide/Tracing%20the%20origin%20of%20the%20coconut%20%28Cocos%20nucifera%20L.%29.pdf |archive-date=June 23, 2021 |access-date=April 22, 2021}}</ref> | |||
[[ | The [[Zulu Social Aid and Pleasure Club]] of [[New Orleans]] traditionally throws hand-decorated coconuts, one of the most valuable [[Mardi Gras]] souvenirs, to parade revelers. The tradition began in the 1910s, and has continued since. In 1987, a "coconut law" was signed by Governor [[Edwin Edwards]] exempting from insurance liability any decorated coconut "handed" from a Zulu float.<ref>{{cite news |first=Mike |last=Scott |title=How the Zulu coconut was saved from extinction |date=February 22, 2017 |url=https://www.nola.com/300/article_c189dcdb-1ec5-5629-84bb-2a2aaec89457.html |newspaper=[[The Times-Picayune/The New Orleans Advocate|The Times-Picayune]] |access-date=May 25, 2020}}</ref> | ||
The coconut is used as a target and prize in the traditional British fairground game [[coconut shy]]. The player buys some small balls which are thrown at coconuts balanced on sticks. The aim is to knock a coconut off the stand and win it.<ref name="harries">{{cite journal |last=Harries |first=Hugh C. |year=2004 |title=Fun Made The Fair Coconut Shy|url=http://www.palms.org/palmsjournal/2004/vol48n2p77-82.pdf |journal=Palms |publisher=[[International Palm Society]] |volume=48 |issue=2 |pages=77–82 |access-date=May 8, 2012 |archive-date=July 23, 2012 |archive-url=https://web.archive.org/web/20120723105002/http://www.palms.org/palmsjournal/2004/vol48n2p77-82.pdf}}</ref> | |||
It was the main food of adherents of the now discontinued Vietnamese [[Coconut Religion]], ''Đạo Dừa''.<ref name="Coconut 1">{{cite web |title=Coconut religion |url=http://vinhthong.info/FAQ/faq_luomlat_CoconutReligion_ConPhung.html |publisher=Vinhthong |archive-url=https://web.archive.org/web/20131022095309/http://vinhthong.info/FAQ/faq_luomlat_CoconutReligion_ConPhung.html |archive-date=October 22, 2013 |access-date=May 25, 2013}}</ref> | |||
=== Myths and legends === | |||
Some South Asian, Southeast Asian, and Pacific Ocean cultures have [[origin myth]]s in which the coconut plays the main role. In the [[Hainuwele]] myth from [[Maluku Islands|Maluku]], a girl emerges from the blossom of a coconut tree.<ref>{{cite book |url=http://www.oxfordreference.com/view/10.1093/acref/9780195156690.001.0001/acref-9780195156690-e-657 |title=Hainuwele – Oxford Reference |isbn=978-0-19-515669-0 |url-status=live |archive-url=https://web.archive.org/web/20130928002738/http://www.oxfordreference.com/view/10.1093/acref/9780195156690.001.0001/acref-9780195156690-e-657 |archive-date=September 28, 2013 |last=Leeming |first=David |date=November 17, 2005 |publisher=[[Oxford University Press]] }}</ref> In [[Maldivian folklore]], one of the main myths of origin reflects the dependence of the [[Maldivians]] on the coconut tree.<ref name="NIAS Press">[[Xavier Romero-Frias|Romero-Frias, Xavier]] (2012) ''Folk tales of the Maldives'', [http://www.niaspress.dk/catalogue2011/Catalogue_2012.pdf NIAS Press] {{webarchive|url=https://web.archive.org/web/20130528131620/http://www.niaspress.dk/catalogue2011/Catalogue_2012.pdf |date=May 28, 2013 }}, {{ISBN|978-87-7694-104-8|978-8776941055}}</ref> In the story of [[Sina and the Eel]], the origin of the coconut is related as the beautiful woman Sina burying an eel, which eventually became the first coconut.<ref>{{cite web |url=http://www.livingheritage.org.nz/schools/secondary/marcellin/samoa/eel.php |archive-url=https://web.archive.org/web/20121029035259/http://www.livingheritage.org.nz/schools/secondary/marcellin/samoa/eel.php |title=Living Heritage – Marcellin College – Sina and the Eel |archive-date=October 29, 2012 |website=www.livingheritage.org.nz}}</ref> | |||
According to urban legend, [[death by coconut|more deaths are caused by falling coconuts than by sharks]] annually, though the truth is actually the reverse.<ref name="flmnh">{{cite web |title=International Shark Attack File |url=http://www.shark.org.au/quotes.html |publisher=Shark Research Institute |archive-url=https://web.archive.org/web/20130729014133/http://shark.org.au/quotes.html |archive-date=July 29, 2013 |access-date=January 9, 2014}}</ref> | |||
=== | === Early history === | ||
Literary evidence from the ''[[Ramayana]]'' and [[History of Sri Lanka|Sri Lankan chronicles]] indicates that the coconut was present in the [[Indian subcontinent]] before the 1st century BCE.<ref name="BlenchSpriggs1998">{{cite book |last1=Blench |first1=Roger |url=https://books.google.com/books?id=DWMHhfXxLaIC&q=coconut&pg=PA396 |title=Archaeology and Language: Correlating archaeological and linguistic hypotheses |author2=Matthew Spriggs |publisher=Routledge |year=1998 |isbn=978-0-415-11761-6 |page=396}}</ref> The earliest direct description is given by [[Cosmas Indicopleustes]] in his ''[[Topographia Christiana]]'' written around 545, where the coconut is called "the great nut of India".<ref name="Rosengarten">Rosengarten, Frederic Jr. (2004). ''The Book of Edible Nuts''. Dover Publications. pp. [https://books.google.com/books?id=7CK8LFCcvtcC&pg=PA65 65–93]. {{ISBN|978-0-486-43499-5}}.</ref> Another early mention is the "[[One Thousand and One Nights]]" story of [[Sinbad the Sailor]], who bought and sold a coconut during his fifth voyage.<ref>{{cite web |date=November 2, 2009 |title=The Fifth Voyage of Sindbad the Seaman – The Arabian Nights – The Thousand and One Nights – Sir Richard Burton translator |url=http://classiclit.about.com/library/bl-etexts/arabian/bl-arabian-5sindbad.htm |url-status=live |archive-url=https://web.archive.org/web/20111225203730/http://classiclit.about.com/library/bl-etexts/arabian/bl-arabian-5sindbad.htm |archive-date=December 25, 2011 |access-date=February 14, 2012 |publisher=Classiclit.about.com }}</ref> | |||
In March 1521, [[Antonio Pigafetta]] described the coconut in his journal in Italian with the word "''cocho''", plural "''cochi''". This followed the first European crossing of the Pacific Ocean during the [[Ferdinand Magellan|Magellan]] [[circumnavigation]]. He explained how at Guam "they eat coconuts" ("''mangiano cochi''") and that the natives there "anoint the body and the hair with coconut and [[sesame seed|beniseed]] oil" ("''ongieno el corpo et li capili co oleo de cocho et de giongioli''").<ref name="ap">{{cite book |last=Pigafetta |first=Antonio |url=https://books.google.com/books?id=283tAAAAMAAJ&q=cocho&pg=PA100 |title=Magellan's Voyage Around the World, Volume 1 (translated by James Alexander Robertson) |publisher=[[Arthur H. Clark Company]] |year=1906 |pages=64–100}}</ref> | |||
== See also == | |||
* [[Domesticated plants and animals of Austronesia]] | * [[Domesticated plants and animals of Austronesia]] | ||
* [[Central Plantation Crops Research Institute]] | * [[Central Plantation Crops Research Institute]] | ||
* [[Coconut production in Kerala]] | * [[Coconut production in Kerala]] | ||
* [[Coir Board of India]] | * [[Coir Board of India]] | ||
==References== | == References == | ||
{{reflist}} | |||
== | === Sources === | ||
* {{cite book | * {{cite book |ref=Grimwood |last1=Grimwood |first1=B. E. |last2=Ashman |first2=F. |year=1975 |title=Coconut Palm Products: Their Processing in Developing Countries |publisher=[[Food and Agriculture Organization]] |url=https://books.google.com/books?id=fY5hLeJ-WW4C&pg=PA1 |isbn=978-92-5-100853-9}} | ||
| last1= | |||
| year= | == External links == | ||
| title= Coconut | |||
| publisher=[[ | |||
| url= https://books.google.com/books?id= | |||
| isbn=978-92- | |||
{{Commons category multi|Coconuts||Cocos nucifera}} | {{Commons category multi|Coconuts||Cocos nucifera}} | ||
{{Wikispecies|Cocos nucifera}} | {{Wikispecies|Cocos nucifera}} | ||
* {{cookbook-inline|Coconut}} | * {{cookbook-inline|Coconut}} | ||
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[[Category:Flora of Florida]] | [[Category:Flora of Florida]] | ||
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[[Category:Pre-Columbian | [[Category:Pre-Columbian transoceanic contact theories]] | ||
[[Category:Oil seeds]] | [[Category:Oil seeds]] | ||
[[Category:Drupes]] | [[Category:Drupes]] | ||
Latest revision as of 18:53, 28 December 2025
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The coconut (Cocos nucifera) is a member of the palm family (Arecaceae) and the only living species of the genus Cocos.[1] The term "coconut" (or the archaic "cocoanut")[2] can denote the whole coconut palm tree or the large hard fruit. Originally native to Central Indo-Pacific, they are ubiquitous in coastal tropical regions.
The coconut tree provides food, fuel, cosmetics, folk medicine and building materials. The inner flesh of the mature fruit forms a regular part of the diets of many people in the tropics and subtropics. Coconut endosperm contains a large quantity of a liquid, "coconut water". Mature coconuts can be processed for oil and coconut milk from the flesh, charcoal from the hard shell, and coir from the fibrous husk. Dried coconut flesh is called copra, and the oil and milk derived from it are commonly used in cooking and in soaps and cosmetics. Sweet coconut sap can be made into drinks or fermented into palm wine or coconut vinegar. The hard shells, fibrous husks and long pinnate leaves are used to make a products for furnishing and decoration.
The coconut has cultural and religious significance for Austronesian peoples, appearing in their mythologies, songs, and oral traditions. It has religious significance in South Asian cultures, where it is used in Hindu rituals including weddings and worship.
The species evolved in the central Indo-Pacific. It was domesticated by Austronesian peoples in Island Southeast Asia and spread during the Neolithic via their seaborne migrations as far east as the Pacific Islands, and as far west as Madagascar. The species played a critical role in their long sea voyages by providing a portable source of food and water, as well as building materials for outrigger boats. Coconuts were spread much later along the coasts of the Indian and Atlantic Oceans by South Asian, Arab, and from the 16th century by European sailors. Based on these introductions, the species can be divided into Pacific and Indo-Atlantic types. The Indo-Atlantic type was introduced to the Americas during the colonial era in the Columbian exchange, while Austronesian sailors appear to have introduced Pacific coconuts to Panama in pre-Columbian times.
Trees can grow up to Script error: No such module "convert". tall and can yield up to 75 fruits per year, though fewer than 30 is more typical. They are intolerant to cold and prefer copious precipitation and full sunlight. Many insect pests and diseases affect commercial production. In 2023, world production of coconuts was 65 million tonnes, with 73% of the total produced by Indonesia, India, and the Philippines.
Description
Cocos nucifera is a large palm, growing up to Script error: No such module "convert". tall, with pinnate leaves Script error: No such module "convert". long, and pinnae Script error: No such module "convert". long; old leaves break away cleanly, leaving the trunk smooth.[3] On fertile soil, a tall coconut palm tree produces around 80 fruits per year; new varieties may be able to yield as many as 150 per year.[4] In India, average production is over 8,000 nuts per hectare per year.[5] Tall varieties produce their first fruit in 6 to 10 years, and live for 60 to 100 years; dwarf varieties become productive more quickly, but have a shorter lifespan.[6]
Inflorescence
The coconut is monoecious, meaning that both male and female flowers grow on the same tree, in its case in the same inflorescence.[7][8] It is possible that the species in addition occasionally has bisexual flowers.[9] The female flower is much larger than the male flower.[8] Mature trees grow continuously, producing leaves, flowers, and fruit all year round. It takes some 14 months for each flower primordium to develop into an inflorescence, botanically a spadix inside a sheathing spathe. A healthy tree can produce up to 15 inflorescences per year, staggered so that there is always a mature one with others in different stages of development.[10][11] It takes 11 months from the opening of the female flowers to the time of harvest.[12] Coconut palms are largely cross-pollinated, although most dwarf varieties are self-pollinating.[13]
Fruit
Botanically, the coconut fruit is a drupe, not a true nut.[14] Like other fruits, it has three layers: the exocarp, mesocarp, and endocarp. The exocarp is the glossy outer skin, usually yellow-green to yellow-brown in color. The mesocarp is composed of a fiber, called coir, which has many traditional and commercial uses. The exocarp and the mesocarp make up the "husk" of the coconut, while the endocarp makes up the hard coconut "shell". The endocarp is around Script error: No such module "convert". thick and has three distinctive germination pores (micropyles) on the distal end. Two of the pores are plugged (the "eyes"), while one is functional.[15][16]
The interior of the endocarp is hollow and is lined with a thin brown seed coat, some Script error: No such module "convert". thick. The endocarp is initially filled with a liquid endosperm (the coconut water). The liquid contains many free cell nuclei (it is multinucleate) dividing by mitosis, without cell boundaries. As development continues, cellular layers of endosperm deposit along the walls of the endocarp up to Script error: No such module "convert". thick, starting at the far end. They eventually form the edible solid endosperm ("coconut meat") which hardens over time. The small cylindrical embryo is embedded in the solid endosperm directly below the functional pore. During germination, the embryo pushes out of the functional pore and forms a haustorium inside the central cavity. This absorbs the solid endosperm to nourish the seedling.[15][17][18]
The fruits have two distinctive forms. Wild niu kafa coconuts feature an elongated triangular fruit with a thicker husk and a smaller amount of endosperm. These make the fruits more buoyant, ideal for ocean dispersal. Domesticated niu vai Pacific coconuts are rounded in shape with a thinner husk, more endosperm, and more coconut water.[19][20][13]
| Component | Mass/kg | ||
|---|---|---|---|
| Husk | 2.033 | ||
| Nut | 1.125 | ||
| of which | Shell | 0.359 | |
| Juice | 0.492 | ||
| Meat | 0.477 | ||
| Total | 3.158 | ||
A full-sized fruit weighs about Script error: No such module "convert". depending on variety.[22][23] The 21st century Bido variety from Indonesia averages Script error: No such module "convert". per fruit.[21]
Coconuts are exported without husks; de-husked nuts from Côte d'Ivoire average around 575 grams, while de-husked nuts from the Dominican Republic average nearly 700 grams.[24] Coconuts sold domestically in coconut-producing countries are typically not de-husked. Immature coconuts (6 to 8 months from flowering) are sold for coconut water and softer jelly-like coconut meat (known as "green coconuts", "young coconuts", or "water coconuts"), where the original coloration of the fruit is more pleasing.[22][23]
Whole mature coconuts (11 to 13 months from flowering) sold for export, however, typically have the husk removed to reduce weight and volume for transport. This results in the naked coconut "shell" with three pores, remnants of the three carpels of the flower, more familiar in countries where coconuts are not grown locally. De-husked coconuts are easier for consumers to open, but have a shorter postharvest storage life of around two to three weeks at temperatures of Script error: No such module "convert". or up to 2 months at Script error: No such module "convert".. In comparison, mature coconuts with a husk can be stored for three to five months at room temperature.[22][23]
Roots
The palm tree has neither a taproot nor root hairs, but a fibrous root system.[7] This consists of many thin roots that grow outward from the plant near the surface. Only a few penetrate deep into the soil for stability. This is known as a fibrous or adventitious root system, and is a characteristic of grass species. 2,000–4,000 adventitious roots may grow, each about Script error: No such module "convert". in diameter. Decayed roots are replaced regularly as the tree grows new ones.[25]
Taxonomy
Taxonomic history
The Swedish botanist and taxonomist Carl Linnaeus formally described the species Cocos nucifera in his book Species Plantarum in 1753.[26] The name is accepted by botanists.[1] In 1768, in his book The Gardeners Dictionary, the English botanist Philip Miller redescribed the plant as Palma cocos, a name treated as a synonym.[27][1] In 1891, the German botanist Otto Kuntze gave it the name Calappa nucifera in his Revisio Generum Plantarum, also treated as a synonym.[28][1]
Etymology
The generic name Cocos, and the common name, is derived from the 16th-century Portuguese word coco, meaning 'head' or 'skull' after the three indentations on the coconut shell that give an impression of a face.[29][30][31][32] This apparently came from encounters in 1521 by Portuguese and Spanish explorers with Pacific Islanders, when the coconut shell reminded them of ghosts in Portuguese folklore called coco or côca.[32][33] In the West, the fruit was originally called nux indica, a name used by Marco Polo in 1280 while in Sumatra. His term is a translation from the Arabic of the time, where it was called جوز هندي jawz hindī, "Indian nut".[6] Thenga, its Tamil/Malayalam name, was used in the detailed description of coconut found in Itinerario by Ludovico di Varthema published in 1510 and in the later Hortus Indicus Malabaricus.[34]
The specific name nucifera means "nut-bearing", from the Latin words nux (nut) and fera (bearing).[35]
Origins
Fossil history
The vast majority of Cocos-like fossils have been recovered from only two regions in the world: New Zealand and west-central India. However, Cocos-like fossils are still putative, as they are difficult to identify.[36] The earliest Cocos-like fossil to be found was C. zeylandica, a fossil species with small fruits, around Script error: No such module "convert". × Script error: No such module "convert". in size, from the Miocene (~23 to 5.3 million years ago) of New Zealand. Since then, numerous other fossils of similar fruits of uncertain affinity have been found in New Zealand from the Eocene, Oligocene, and possibly the Holocene.[36][37] In the Deccan Traps of west-central India, numerous fossils of Cocos-like fruits, leaves, and stems have been found. They include morphotaxa like Palmoxylon sundaran, Palmoxylon insignae, and Palmocarpon cocoides. Cocos-like fossils of fruits include Cocos intertrappeansis, Cocos pantii, and Cocos sahnii. Some have been tentatively identified as modern C. nucifera. These include two specimens named C. palaeonucifera and C. binoriensis, both dated by their authors to the Maastrichtian–Danian of the early Tertiary (70 to 62 million years ago). C. binoriensis has been claimed to be the earliest known fossil of C. nucifera.[38][36][39]
Only two other regions have reported Cocos-like fossils, namely Australia and Colombia. In Australia, a Cocos-like fossil fruit, measuring Script error: No such module "convert"., was recovered from the Chinchilla Sand Formation dated to the latest Pliocene or basal Pleistocene. Rigby (1995) assigned them to modern Cocos nucifera based on its size.[38][36] In Colombia, a single Cocos-like fruit was recovered from the middle to late Paleocene Cerrejón Formation. The fruit, however, was compacted in the fossilization process and it was not possible to determine if it had the diagnostic three pores that characterize members of the tribe Cocoseae. Nevertheless, one study assigned it to Cocos based on the size and the ridged shape of the fruit.[40]
Phylogeny
A 2016 molecular phylogenomic analysis of the palms places the genus Cocos among the tribe Cocoseae:[41]
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Human dispersal
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Genetic studies identify the coconut's center of origin as the Central Indo-Pacific, where it has its greatest genetic diversity.[45][25][46][47] Its cultivation and spread was closely tied to migrations of the Austronesian peoples who carried coconuts to the islands they settled.[46][47][48][49] Linguistic, archaeological, and genetic evidence all points to domestication of Pacific coconuts by Austronesians in Southeast Asia during the Austronesian expansion (c. 3000 to 1500 BCE).[50][47][46][51] Drift models based on wind and ocean currents show that coconuts could not have drifted across the Pacific unaided,[52] implying that dispersal was human-assisted.[53]
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Coconuts are divided into two subpopulations, a Pacific group from Island Southeast Asia and an Indo-Atlantic group from the south of the Indian subcontinent. The Pacific group is clearly domesticated, with dwarf habit, self-pollination, and niu vai fruits with large endosperm-to-husk ratios. The distribution of Pacific coconuts corresponds to regions settled by Austronesian voyagers, especially Madagascar. The island's coconuts show genetic admixture between the two subpopulations, indicating that Pacific coconuts interbred with Indo-Atlantic coconuts there.[47][48] Although archaeological remains from 1000 to 500 BCE suggest that Indo-Atlantic coconuts were later independently cultivated by Dravidian peoples, only Pacific coconuts show clear domestication traits like dwarf habits, self-pollination, and rounded fruits. Indo-Atlantic coconuts, in contrast, have the ancestral traits of tall habits and elongated triangular fruits.[50][47][46][51]
Genetic studies have confirmed pre-Columbian populations of coconuts in Panama. However, it is not native and displays a genetic bottleneck resulting from a founder effect. Coconuts in the Americas are most closely related to those in the Philippines, indicating that the coconuts were not introduced naturally, such as by sea currents, but by early Austronesian sailors to the Americas from at least 4250 BCE.[46][49][52] During the colonial era, Pacific coconuts were further introduced to Mexico from the Spanish East Indies via the Manila galleons, starting in the 16th century. In contrast, Indo-Atlantic coconuts were spread by Arab and Persian traders into the East African coast. Indo-Atlantic coconuts were introduced into the Atlantic Ocean by Portuguese ships from colonies in India and Sri Lanka, again starting in the 16th century: first to coastal West Africa, and then to the Caribbean and Brazil.[47]
Domestication
Coconuts can be broadly divided into two fruit types – the ancestral niu kafa form with a thick-husked, angular fruit, and the niu vai form with a thin-husked, spherical fruit with a higher proportion of endosperm. The terms are Samoan.[47][19][54]
The niu kafa form is the wild ancestral type, with thick husks to protect the seed, and an angular, highly ridged shape to promote buoyancy during ocean dispersal. It is the dominant form in the Indo-Atlantic coconuts.[19][47] However, they may have been selected to some extent for thicker husks for coir production, which was important in Austronesian material culture as a source for cordage in building houses and boats.[50]
- Two major fruit types
-
The niu kafa form of wild and Indo-Atlantic coconuts, possibly selected for more coir for houses and boats
The niu vai form is the domesticated form dominant in Pacific coconuts. They were artificially selected by Austronesians for their larger endosperm-to-husk ratio and higher coconut water content, making them more useful as food and water reserves for sea voyages. The decreased buoyancy and increased fragility of this spherical, thin-husked fruit did not matter for a species that had started to be dispersed by humans and grown in plantations.[19][20] Niu vai endocarp fragments have been recovered in archaeological sites in the St. Matthias Islands of the Bismarck Archipelago. The fragments are dated to approximately 1000 BCE, suggesting that cultivation and artificial selection of coconuts were already practiced by the Austronesian Lapita people.[50]
Coconuts can be broadly divided into two general types based on habit: the "Tall" (var. typica) and "Dwarf" (var. nana) varieties.[55] The two groups are genetically distinct, with the dwarf variety showing a greater degree of artificial selection for ornamental traits and for early germination and fruiting.[54][56] The tall variety is outcrossing while dwarf palms are self-pollinating, which has led to a much greater degree of genetic diversity within the tall group.[57]
The dwarf coconut cultivars are fully domesticated, unlike the more diverse tall cultivars.[58][57] Dwarf coconuts share three genetic markers out of thirteen (rare in tall cultivars), making it likely that they originate from a single domesticated population. Philippine and Malayan dwarf coconuts diverged early into two distinct types. They usually remain genetically isolated when introduced to new regions. Numerous other dwarf cultivars developed after such introductions, hybridizing with tall cultivars. The origin of dwarf varieties is Southeast Asia, which contain the tall cultivars genetically closest to dwarf coconuts.[47][59][58][57]
Genome sequencing of tall and dwarf varieties reveals that they diverged 2 to 8 million years ago and that the dwarf variety arose through alterations in genes for the metabolism of the plant hormone gibberellin.[60]
Another ancestral variety is the niu leka of Polynesia (the "Compact Dwarfs"). Although it resembles dwarf coconuts (including slow growth), it is genetically distinct and was independently domesticated, likely in Tonga. Other cultivars of niu leka may exist in other islands of the Pacific, and some are probably descendants of advanced crosses between Compact Dwarfs and Southeast Asian Dwarf types.[59][58]
Distribution and habitat
Coconuts have a nearly cosmopolitan distribution due to human cultivation and dispersal. However, their original distribution was in the Central Indo-Pacific, in the regions of Maritime Southeast Asia and Melanesia.[45]
The coconut palm thrives on sandy soils and is highly tolerant of salinity. It prefers areas with abundant sunlight and regular rainfall of between 1500 mm and 2500 mm per year. It prefers humidity above 60%. If rainfall is less than this, it can survive if its roots can reach the soil water table, but it cannot tolerate waterlogging. It grows from sea level to an altitude of 600 metres in the tropics. It can tolerate a dry season of one month on sandy soils inland, and as much as three months on heavier soils, but the soil must be free-draining. It grows on soils with a pH of 4.5 to 8 (the latter on coral atolls), but prefers a range of 5.5 to 7. Growth is seriously limited by shade. It can resist hurricane-strength winds provided it has developed a good root system.[61]
Wild coconuts are restricted to coastal areas in sandy, saline soils. The fruit is adapted for ocean dispersal. Coconuts could not reach inland locations without human intervention to carry seednuts and plant seedlings.[62]
Cultivation
Coconut palms are normally cultivated in hot and wet tropical climates. They need year-round warmth and moisture to grow well and fruit. Coconut palms are hard to establish in dry climates, and cannot grow there without frequent irrigation. In drought conditions, the new leaves do not open well, older leaves may become desiccated, and fruit may be shed.[61]
The extent of cultivation in the tropics is threatening a number of habitats, such as mangroves; an example of such damage to an ecoregion is in the Petenes mangroves of the Yucatán.[63] Uniquely among trees, coconut trees can be irrigated with sea water.[64]
Pests and diseases
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Coconuts are susceptible to the phytoplasma disease, lethal yellowing. Yellowing diseases affect plantations in Africa, India, Mexico, the Caribbean and the Pacific Region.[65]
The coconut palm is damaged by the larvae of many Lepidoptera (butterfly and moth) species which feed on it, including the African armyworm (Spodoptera exempta) and Batrachedra spp.: B. arenosella, B. atriloqua (feeds exclusively on C. nucifera), B. mathesoni (feeds exclusively on C. nucifera), and B. nuciferae.[66]
The coconut leaf beetle Brontispa longissima feeds on young leaves, and damages both seedlings and mature coconut palms. In 2007, the Philippines imposed a quarantine in Metro Manila and 26 provinces to stop the spread of the pest and protect the Philippine coconut industry managed by some 3.5 million farmers.[67]
The fruit may be damaged by eriophyid coconut mites (Aceria guerreronis). This mite infests coconut plantations, and can cause economic damage up to 60% of coconut production.[68] The immature seeds are infested and damaged by larvae.[69] Chemical control is possible, but since it needs to be repeated frequently it is impracticable on grounds of cost, environmental harm, and pesticide residues in coconut meat and coconut water.[70]
Cultivars
Coconut has many commercial and traditional cultivars. They can be sorted mainly into tall, dwarf, and hybrid cultivars (hybrids between talls and dwarfs).[71] Varieties are often regional, such as Ceylon Tall, Jamaica Tall, Java Tall, and Malayan Tall.[6]
Dwarf varieties of Pacific coconuts have been cultivated by Austronesian peoples since ancient times. These varieties were selected for slower growth, sweeter coconut water, and often brightly colored fruits.[59] Dwarf varieties include Dwarf Green and Dwarf Orange.[6]
Varieties have been selected for diverse traits: for example, King coconut is a Sri Lankan variety with a relatively low sugar content, while Macapuno has soft jelly-like flesh that fills the whole central cavity; it is used to make sweet desserts.[72][73]
Maypan is an F1 hybrid bred in Jamaica in the 1970s to resist lethal yellowing.[74][75] However, Maypan resistance soon began to fail, possibly as early as the 1980s, and certainly by the 2000s.[76][77] Some other coconut varieties have natural resistance to lethal yellowing with alleles at microsatellites, with Vanuatu tall and Sri-Lanka green dwarf as the most resistant cultivars, while West African tall is especially susceptible.[78][79][80]
Breeding
Conventional plant breeding is of limited use with coconut because there is no wild coconut species to supply additional genetic diversity; the generation time is long; there is much heterozygosity; artificial pollination to conduct crosses produces only a few seeds; and vegetative reproduction (cloning) is unreliable. Coconut breeding objectives can include copra content, production of female flowers, oil content, aroma of coconut meat and water, softness and sweetness of endosperm, drought tolerance, resistance to root wilt, and resistance to eriophorid mites.[81]
Hybrids provide higher precocity and productivity with the number of fruits than other breeds, but they produce fruits with low market acceptance for the water quality of the fruit. Intravarietal crosses in Dwarf coconut (dwarf coconuts bred together) have been tested to provide better water quality for the coconut water market than the hybrids.[82]
Harvesting
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The two most common harvesting methods are by climbing[83] and by using poles.[84]
Climbing is the more widespread, but it is more dangerous and requires skilled workers.[83] Manually climbing trees is traditional in most countries and requires a posture that exerts pressure on the trunk with the feet. Climbers employed on coconut plantations often develop musculoskeletal disorders and risk injury or death from falling.[85][86][87] For safety, coconut climbers in the Philippines and Guam use a bolo knife tied with a rope to the waist to cut grooves at regular intervals on the coconut trunks. This makes the trunk of the tree more like a ladder, though it reduces the value of timber recovered from the trees and can admit infection.[88][83][89] Other methods to make climbing easier include using a system of pulleys and ropes; using pieces of vine, rope, or cloth tied to both hands or feet; using spikes attached to the feet or legs; or attaching coconut husks to the trunk with ropes.[90]
The pole method uses a long pole with a cutting device at the end. In the Philippines, the traditional tool is called the halabas and is made from a long bamboo pole with a sickle-like blade at its tip. Though safer and faster than climbing, it does not allow workers to examine and clean the crown of coconuts for pests and diseases.[84]
Modern methods use hydraulic elevators mounted on tractors or ladders.[91] Mechanical coconut climbing devices and robots have been developed in India, Sri Lanka, and Malaysia.[92][93][94][90] The Coconut Maturity Detection Project uses imaging and machine learning to identify mature coconut bunches ready for harvesting.[95]
A system of bamboo bridges and ladders directly connecting the tree canopies is used in the Philippines for coconut plantations that harvest coconut sap (not fruits) for coconut vinegar and palm wine production.[96][91] In other areas, as in Papua New Guinea, coconuts are simply collected when they fall to the ground.[83]
Some coconut farmers in Thailand and Malaysia use southern pig-tailed macaques to harvest coconuts.[97] Thailand has been raising and training them to pick coconuts for around 400 years.[98][99][100] Training schools for pig-tailed macaques still exist in southern Thailand and in the Malaysian state of Kelantan.[101] People for the Ethical Treatment of Animals (PETA) objected to the Thailand practice in 2019.[91]
| Script error: No such module "flag". | 18.0 |
| Script error: No such module "flag". | 14.9 |
| Script error: No such module "flag". | 14.2 |
| Script error: No such module "flag". | 2.9 |
| Script error: No such module "flag". | 2.1 |
| World | 64.7 |
|---|---|
| Source: FAOSTAT of the United NationsScript error: No such module "Check for unknown parameters".[102] | |
Production
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In 2023, world production of coconuts in shells was 65 million tonnes, led by Indonesia, India, and the Philippines, with 73% of the total when combined (table).
In India, four southern states account for most of India's total production: Tamil Nadu, Karnataka, Kerala, and Andhra Pradesh.[103] Though Kerala has the largest number of coconut trees, Tamil Nadu is the most productive per hectare.[104] The coconut is the official state tree of Kerala, whose name in the local language Malayalam means "coconut land".[105]
The main coconut-producing area in the Middle East is the Dhofar region of Oman.[106] The ancient coconut groves of Dhofar were mentioned by the medieval Moroccan traveller Ibn Battuta in his book The Rihla.[107]
Sri Lanka is the fourth-largest producer, and is the second-largest producer of coconut oil and copra, accounting for 15% of global production. The production of coconuts is the main source of Sri Lanka economy, with 12% of cultivated land and 409,244 hectares used for coconut growing (2017). Sri Lanka established its Coconut Development Authority and Coconut Cultivation Board and Coconut Research Institute in the early British Ceylon period.[108]
Health concerns
Cardiovascular disease
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As a rich source of lauric acid and other saturated fats, coconut oil is a dietary risk factor for cardiovascular diseases by elevating blood levels of LDL cholesterol.[109][110] Many national clinical associations recommend limiting consumption of coconut oil products, replacing them instead with foods containing unsaturated fats.[109][110][111]
Allergens
Proteins from coconut in food may cause allergic reactions, including anaphylaxis, in some people.[112] In the United States, the Food and Drug Administration has declared that coconut must be disclosed as an ingredient on food package labels as a "tree nut" with potential allergenicity.[113]
Cocamidopropyl betaine (CAPB) is a surfactant manufactured from coconut oil and used in personal hygiene products and cosmetics, such as shampoos, liquid soaps, cleansers and antiseptics. CAPB may cause mild skin irritation,[114] but allergic reactions to CAPB are rare;[115] they are probably related to impurities rendered during the manufacturing process (which include amidoamine and dimethylaminopropylamine) rather than CAPB itself.[114]
Uses
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The coconut palm is grown throughout the tropics for decoration, as well as for its culinary and nonculinary uses; virtually every part of the coconut palm is used by humans in some manner and has significant economic value. The coconut's versatility is noted in some of its names: in Sanskrit, it is kalpa vriksha ("the tree of the necessities of life"), while in Malay, it is pokok seribu guna ("the tree of a thousand uses"), and in the Philippines, it is called the "tree of life".[116] It is one of the most useful trees in the world.[17]
Nutrition
Template:Nutritional value Raw coconut meat is 47% water, 33% fat, 15% carbohydrates, and 3% protein (table). In a reference amount of Script error: No such module "convert"., raw coconut flesh supplies 350 calories of food energy, and is a rich source (20% or more of the Daily Value, DV) of manganese (65% DV) and copper (48% DV), with various other dietary minerals in moderate amounts (10-18% DV; table). It is a poor source of vitamins. Raw coconut meat has a high content of saturated fatty acids (30% of total fats), with lauric acid as the main saturated fat (15% of total; USDA source in table).
Culinary
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The many culinary uses of coconuts are largely based on the edible white, fleshy part of the seed (the endosperm), known as "coconut meat".[117] The meat of immature coconuts can be eaten as it is or cooked in pastries. Mature coconut meat is tough and is processed before consumption, being made into products like coconut milk,[117][118][119][120][121] "coconut chips"[120] or grated and dehydrated as "desiccated coconut".[122][123][124]
Coconut milk, used for cooking many dishes, is pressed from coconut meat. It can be diluted to create coconut milk beverages such as milk substitutes.[125][92] Coconut milk powder, a protein-rich powder, can be processed from coconut milk.[126] Coconut milk and coconut cream extracted from grated coconut can be added to desserts and savory dishes, or used in curries and stews.[127][128] Products made from thickened coconut milk with sugar and eggs, like coconut jam and coconut custard, are widespread in Southeast Asia.[129][130][131] Coconut oil is used for frying and cooking.[127][132]
Coconut water can be drunk fresh or used in cooking.[133][134] It can be fermented to produce a jelly-like dessert known as nata de coco.[135] Coconut vinegar, made from fermented coconut water or sap, is used extensively in Southeast Asian and Goan cuisine.[118]
Coconut sap, fresh or fermented, is drunk as toddy or tubâ in the Philippines. When left to ferment on its own, it becomes palm wine. Palm wine is distilled to produce arrack.[136] The sap can be reduced by boiling to create a sweet syrup, or reduced further to yield coconut sugar. A young, well-maintained tree can produce around Script error: No such module "convert". of toddy per year, while a 40-year-old tree may yield around Script error: No such module "convert"..[137]
Oil
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| Script error: No such module "flag". | 1,358,900 |
| Script error: No such module "flag". | 666,000 |
| Script error: No such module "flag". | 360,000 |
| Script error: No such module "flag". | 182,000 |
| Script error: No such module "flag". | 132,300 |
| Script error: No such module "flag". | 84,015 |
| World | 3,162,279 |
|---|---|
| Source: FAOSTATScript error: No such module "Check for unknown parameters".[138] | |
In 2022, world production of processed coconut oil was 3.2 million tonnes, led by the Philippines with 43% of the total, and Indonesia and India as secondary producers (table).
Coconut oil is used in cooking, especially for frying. It can be used in liquid form like other vegetable oils, or in solid form like butter or lard.[139][140][141] Coconut butter is a solidified coconut oil, but the name is also applied to creamed coconut, a specialty product made of coconut milk solids or puréed coconut meat and oil.[117]
Non-food uses
Among the many non-food uses of coconut palms, the husk and shells can be used for fuel or made into charcoal.[142] The husks can serve as flotation devices or as an abrasive.[143] The shell, freed from the husk, and heated on warm ashes, exudes an oily material that is used to soothe dental pains in traditional medicine of Cambodia.[144] Coir fiber from husks is used in ropes, mats, brushes, and sacks, as caulking for boats, and as stuffing for mattresses.[145] It is used in horticulture in potting compost, especially in orchid mix, and to make brooms in Cambodia.[144] Coconut cups were frequently carved with scenes in relief and mounted with precious metals.[146] The leaves provide material for baskets and for roofing thatch; they can be woven into mats, cooking skewers, and kindling arrows. Leaves are woven into small pouches that are filled with rice and cooked to make pusô and ketupat.[147]
Hawaiians hollowed out coconut trunks to form drums, containers, or small canoes. The "branches" (leaf petioles) are strong and flexible enough to make a switch. The use of coconut branches in corporal punishment was revived in the Gilbertese community on Choiseul in the Solomon Islands in 2005.[148] The roots are used to make dye, a mouthwash, and a folk medicine for diarrhea and dysentery.[149] A frayed piece of root can be used as a toothbrush. In Cambodia, the roots are used in traditional medicine.[144] Leftover fiber from coconut oil and coconut milk production, coconut meal, is used as livestock feed. The dried calyx is used as fuel in wood-fired stoves. Coconut water is traditionally used as a growth supplement in plant tissue culture and micropropagation.[150]
In culture
A coconut (Template:Langx) is used in Hindu rituals.[151] Often it is decorated with bright metal foils. It is offered during worship to a Hindu god or goddess. Narali Purnima is celebrated on a full moon day which usually signifies the end of monsoon season in India. The word Narali is derived from Marathi naral, "coconut". Fishermen give an offering of coconut to the sea to celebrate the beginning of a new fishing season,[152] in the hope of bountiful catches. Hindus often initiate any new activity by breaking a coconut to ensure the blessings of the gods. The Hindu goddess of well-being and wealth, Lakshmi, is often shown holding a coconut.[153] They are used in Hindu weddings as a symbol of prosperity.[154] The flowers are used sometimes in wedding ceremonies in Cambodia.[144] The coconut has cultural and religious significance for Austronesian peoples, appearing in their mythologies, songs, and oral traditions.[155][50]
The Zulu Social Aid and Pleasure Club of New Orleans traditionally throws hand-decorated coconuts, one of the most valuable Mardi Gras souvenirs, to parade revelers. The tradition began in the 1910s, and has continued since. In 1987, a "coconut law" was signed by Governor Edwin Edwards exempting from insurance liability any decorated coconut "handed" from a Zulu float.[156]
The coconut is used as a target and prize in the traditional British fairground game coconut shy. The player buys some small balls which are thrown at coconuts balanced on sticks. The aim is to knock a coconut off the stand and win it.[157]
It was the main food of adherents of the now discontinued Vietnamese Coconut Religion, Đạo Dừa.[158]
Myths and legends
Some South Asian, Southeast Asian, and Pacific Ocean cultures have origin myths in which the coconut plays the main role. In the Hainuwele myth from Maluku, a girl emerges from the blossom of a coconut tree.[159] In Maldivian folklore, one of the main myths of origin reflects the dependence of the Maldivians on the coconut tree.[160] In the story of Sina and the Eel, the origin of the coconut is related as the beautiful woman Sina burying an eel, which eventually became the first coconut.[161]
According to urban legend, more deaths are caused by falling coconuts than by sharks annually, though the truth is actually the reverse.[162]
Early history
Literary evidence from the Ramayana and Sri Lankan chronicles indicates that the coconut was present in the Indian subcontinent before the 1st century BCE.[163] The earliest direct description is given by Cosmas Indicopleustes in his Topographia Christiana written around 545, where the coconut is called "the great nut of India".[164] Another early mention is the "One Thousand and One Nights" story of Sinbad the Sailor, who bought and sold a coconut during his fifth voyage.[165]
In March 1521, Antonio Pigafetta described the coconut in his journal in Italian with the word "cocho", plural "cochi". This followed the first European crossing of the Pacific Ocean during the Magellan circumnavigation. He explained how at Guam "they eat coconuts" ("mangiano cochi") and that the natives there "anoint the body and the hair with coconut and beniseed oil" ("ongieno el corpo et li capili co oleo de cocho et de giongioli").[166]
See also
- Domesticated plants and animals of Austronesia
- Central Plantation Crops Research Institute
- Coconut production in Kerala
- Coir Board of India
References
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Sources
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External links
Template:Sister project Template:Sister project
Template:Coconut Template:Culinary wrappings Template:Nuts Template:Non-timber forest products Template:Arecaceae genera Template:Taxonbar Template:Authority control
- Pages with script errors
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- Coconuts
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