Old World babbler

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The Old World babblers or Timaliidae, are a family of mostly Old World passerine birds. They are rather diverse in size and coloration, but are characterised by soft, fluffy plumage. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian subcontinent. The timaliids are one of two unrelated groups of birds known as babblers, the other being the Australasian babblers of the family Pomatostomidae (also known as pseudo-babblers). This family takes its name from the genus Timalia: this generic name was coined by the French zoologist Georges Cuvier in 1821. Although Cuvier did not explain its etymology, it is thought to be based on a Latinized form of an Asian vernacular name (probably Hindu or Indo-Malaysian languages) for these birds, as many species in this family are native to South and Southeast Asia.

Morphological diversity is rather high; most species resemble "warblers", jays or thrushes. This group is among those Old World bird families with the highest number of species still being discovered.

Characteristics

Timaliids are small to medium birds. They have strong legs, and many are quite terrestrial. They typically have generalised bills, similar to those of a thrush or warbler, except for the scimitar babblers which, as their name implies, have strongly decurved bills. Most have predominantly brown plumage, with minimal difference between the sexes, but many more brightly coloured species also exist.[1]

This group is not strongly migratory, and most species have short rounded wings, and a weak flight. They live in lightly wooded or scrubland environments, ranging from swamp to near-desert. They are primarily insectivorous, although many will also take berries, and the larger species will even eat small lizards and other vertebrates.[1]

Typical babblers live in communities of around a dozen birds, jointly defending a territory. Many even breed communally, with a dominant pair building a nest, and the remainder helping to defend and rear their young. Young males remain with the group, while females move away to find a new group, and thus avoid inbreeding. They make nests from twigs, and hide them in dense vegetation.[1]

Taxonomy and systematics

The systematics of Old World babblers have long been contested. During much of the 20th century, the family was used as a "wastebin taxon" for numerous hard-to-place Old World songbirds (such as Picathartidae and Pnoepygidae, as well as the New World species the wrentit). The German ornithologist Ernst Hartert summarized this attitude with the statement that, in the passerines: "Script error: No such module "Lang"." (What one can't place systematically is considered an Old World babbler).[2]

The most obviously misplaced taxa were removed piecemeal towards the end of the last century. Since then, with the aid of DNA sequence data, it has been confirmed that even the remaining group is not monophyletic. Analysis of mtDNA cytochrome b and 12S/16S rRNA data (Cibois 2003a) spread the Timaliidae that were studied across what essentially was a badly resolved polytomy with Old World warblers and white-eyes. As the typical warblers (genus Sylvia) grouped with some presumed timaliids (such as the fulvettas), it was suggested that some Sylviidae should be moved to the Timaliidae.

The phylogenetic relationships between Timaliidae and other families was determined in a molecular phylogenetic study by Tianlong Cai and collaborators that was published in 2019. It is shown in the cladogram below:[3][4]

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The cladogram below shows the relationships between the genera. These were determined in the same study by Cai and collaborators.[3][4] Script error: No such module "Clade".

List of genera

The family as currently constituted includes 58 species divided into the following ten genera:[4]

Image Genus Living species
File:Large Scimitar-babbler - Thailand.jpg Erythrogenys Baker, 1930Script error: No such module "Check for unknown parameters". – scimitar-babblers
File:Streak-breasted Scimitar Babbler.jpg Pomatorhinus Horsfield, 1821Script error: No such module "Check for unknown parameters". – scimitar-babblers
File:Bar-winged Wren-Babbler - Eaglenest - India FJ0A8141 (33475375613).jpg Spelaeornis David & Oustalet, 1877Script error: No such module "Check for unknown parameters". – typical wren-babblers
File:Grey-throated babbler Zuluk East Sikkim Sikkim India 24.05.2015.jpg Stachyris Hodgson, 1844Script error: No such module "Check for unknown parameters".
File:Chestnut-winged Babbler, Danum Valley, Borneo (5836179271).jpg Cyanoderma Salvadori, 1874Script error: No such module "Check for unknown parameters".[5]
File:Tawny bellied warbler 2 by David Raju (cropped).jpg Dumetia Blyth, 1852Script error: No such module "Check for unknown parameters".
File:Pin-striped Tit-babbler Macronus gularis by Dr. Raju Kasambe DSC 5961 (65).jpg Mixornis Blyth, 1842Script error: No such module "Check for unknown parameters". – tit-babblers
File:Macronus striaticeps.jpg Macronus Jardine & Selby, 1835Script error: No such module "Check for unknown parameters". – tit-babblers
File:Chestnut-capped-Babbler (8713230888).jpg Timalia Horsfield, 1821Script error: No such module "Check for unknown parameters".
File:0A2A1562 Bare-headed Laughingthrush.jpg Melanocichla Sharpe, 1883Script error: No such module "Check for unknown parameters".

References

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External links

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