Lufeng Formation

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Template:Short description Script error: No such module "Infobox".Template:Template otherScript error: No such module "check for unknown parameters". The Lufeng Formation (formerly Lower Lufeng Series) is a Lower Jurassic sedimentary rock formation found in Yunnan, China. It has two units: the lower Dull Purplish Beds/Shawan Member are of Hettangian age, and Dark Red Beds/Zhangjia'ao Member are of Sinemurian age.[1] It is known for its fossils of early dinosaurs. The Dull Purplish Beds have yielded the possible therizinosaur Eshanosaurus, the possible theropod Lukousaurus, and the "prosauropods" "Gyposaurus" sinensis, Lufengosaurus, Jingshanosaurus, and Yunnanosaurus. Dinosaurs discovered in the Dark Red Beds include the theropod Sinosaurus triassicus, the "prosauropods" "Gyposaurus", Lufengosaurus, and Yunnanosaurus, indeterminate remains of sauropods, and the early armored dinosaurs Bienosaurus and Tatisaurus.[2]

Paleofauna

Rhynchocephalians

Rhynchocephalians reported from the Lufeng Formation
Genus Species Location Stratigraphic position Material Notes

Clevosaurus[3]

Indeterminate

Yunnan

Partial skulls and jaws.[3] The three named species do not display any autapomorphic characters and should be considered indeterminate within the genus. Only record of rhynchocephalians from Asia.[3]

Crurotarsans

Crurotarsans reported from the Lufeng Formation
Genus Species Location Stratigraphic position Material Notes

Dianchungosaurus[4]

D. lufengensis[4]

Yunnan[4]

Dark Red Beds[4]

Formerly considered an ornithopod dinosaur.

Dianosuchus[5] D. changchiawaensis Lufeng, Yunnan Dark Red Beds
Microchampsa[6] M. scutata Dahuangtian locality Lower Red Beds iIncomplete skeleton consisting of cervical and anterior dorsal vertebrae as well as ribs and three rows of dorsal osteoderms
Phyllodontosuchus[7] P. lufengensis Dawa, Yunnan Dark Red Beds BVP568-L12, a crushed skull A potential non-carnivorous crocodrylomorph
Platyognathus[8] P. hsui Lufeng, Yunnan Zhangjiawa Member IVPP V8266, rostral portion of a skull missing the dorsal bones and with articulated incomplete mandible; CVEB 21301, a nearly complete skull with articulated mandible, articulated with the anterior 17 vertebrae and associated dorsal osteoderms; part of the right scapulocoracoid; the left humerus; the right femur and proximal tibia and fibula The oldest Gobiosuchoidean
Strigosuchus[9] S. licinus Lufeng, Yunnan Zhangjia'ao Member FMNH CUP 2082

Ornithischians

Indeterminate ornithopod remains Yunnan. Dark Red Beds.[2]

Ornithischians reported from the Lufeng Formation
Genus Species Location Stratigraphic position Material Notes

Bienosaurus[4][10]

B. lufengensis[4]

Yunnan[4]

Dark Red Beds[4]

A right "[d]entary with teeth,"[11] with additional cranial fragments such as a partial frontal. These specimens are catalogued as IVPP V 9612. The dentary preserves 11 teeth or roots with two additional empty alveoli.

Tatisaurus[4]

T. oehleri[4]

Yunnan[4]

Dark Red Beds[4]

"Isolated dentary."[11]

Sauropodomorphs

Template:Paleobiota-key-compact

Sauropodomorphs reported from the Lufeng Formation
Genus Species Location Stratigraphic position Material Notes Images

Chuxiongosaurus[12]

C. lufengensis[12]

Yunnan[12]

"Skull[12]

File:Gyposaurus-outline-basicidea.svg
"Gyposaurus" sinensis
File:Jingshanosaurus xinwaensis.png
Jingshanosaurus
File:Lishulong wangi.png
Lishulong
File:Lufengosaurus sketch2.jpg
Lufengosaurus
File:Xingxiulong yueorum.png
Xingxiulong
File:Yunnanosaurus BW.jpg
Yunnanosaurus

Gyposaurus[13]

G. sinensis[13]

Yunnan[13]

  • Dark Red Beds[4]
  • Dull Purplish Beds[14]

"[Two] skeletons, [one] with partial skull, [two] partial skeletons, [three] skull fragments, adult."[15]

Fulengia[4]

F. youngi[4]

Yunnan[4]

  • Dark Red Beds[4]

"Skull."[16]

Jingshanosaurus[14]

J. xinwaensis[14]

Yunnan[14]

  • Dull Purplish Beds[14]

"Complete skeleton with skull, adult."[15]

"Kunmingosaurus"[4]

"K. wusdingensis"[4]

Yunnan[4]

  • Dark Red Beds[4]

nomen nudum

Lishulong[17] L. wangi[17] Yunnan[17] Skull and cervical vertebrae 2–10[17]

Lufengosaurus[13]

L. huenei[13]

Yunnan[13]

"(including Gyposaurus sinensis, L. magnus)"[2]

L. magnus[13]

Yunnan[13]

Tawasaurus[4]

T. minor[4]

Yunnan[4]

  • Dark Red Beds[4]
Xingxiulong X. chengi Yunnan
X. yueorum[18] Yunnan
  • Zhangjiaao Member
 LF2015-NO01, an articulated postcranial skeleton
Yizhousaurus[19] Y. sunae Yunnan
  • Zhangjiaao Member
 Partial skeleton with skull

Yunnanosaurus[13]

Y. huangi[13]

Yunnan[13]

"More than [twenty] partial to complete skeletons, [two] skulls, juvenile to adult."[16]

Y. robustus[13]

Yunnan[13]

Theropods

Theropods reported from the Lower Lufeng Formation
Genus Species Location Stratigraphic position Material Notes Images

Eshanosaurus[14]

E. deguchiianus[14]

Yunnan[14]

  • Dull Purplish Beds[14]

"Dentary."[20]

Possible therizinosaur

File:Diloph sin DB1.jpg
Sinosaurus triassicus

Lukousaurus[13]

L. yini[13]

Yunnan[13]

  • Dark Red Beds[4]
  • Dull Purplish Beds[14]
  • Holotype skull, tooth[21]
  • Three bone fragments[22]

Possible crocodylomorph[23]

Sinosaurus[13]

S. triassicus[13]

Yunnan[13]

  • Dark Red Beds[4]
  • Dull Purplish Beds[14]
  • Maxillary fragments, teeth, and a lower jaw fragment
  • Incomplete skull and other post-cranial fragments

Dilophosaurus sinensis specimen

Now included in Sinosaurus

Panguraptor[24] P. lufengensis[24] Yunnan[24]
  • Dull Purplish Beds[24]
  • Partial skeleton[24]
 A coelophysid

Cynodonts

Cynodonts reported from the Lufeng Formation
Genus Species Location Stratigraphic position Material Notes Images
Bienotherium
  • B. yuannanese
  • B. magnum
 A tritylodontid
File:Morganucodon.svg
Skull of Morganucodon oehleri
Dianzhongia D. longirostrata A tritylodontid
Hadrocodium[25]

H. wui[25]

Yunnan[25]

  • Dark Red Beds
 Skull One of the oldest and smallest mammaliaforms known. Indicates a correlation between the separation of the middle ear bones from the mandible and the expanded brain vault in early mammals.[26]
Lufengia L. delicata A tritylodontid
Morganucodon
  • M. oehleri
  • M. heikuopengensis
 Zhangjiawa Member (M. heikuopengensis)

Shawan Member (M. oehleri)

A morganucodontan
Sinoconodon S. rigneyi Zhangjiawa Member A mammaliamorph closely related to Mammaliaformes
Yunnanodon[27][28]

Y. brevirostre[27]

Yunnan[27]

  • Dark Red Beds
 A tritylodontid

See also

References

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  1. ↑ Luo, Z., and X.-C. Wu. 1994. The small tetrapods of the Lower Lufeng Formation, Yunnan, China; pp. 251–270 in N. C. Fraser and H.-D.Sues (eds.), In the Shadow of the Dinosaurs. Cambridge University Press, New York
  2. ↑ a b c Weishampel, et al. 2004.
  3. ↑ a b c Jones MEH. 2006 The Early Jurassic clevosaurs from China (Diapsida: Lepidosauria). New Mex. Museum Nat. Hist. Sci. Bull. 37, 548–562.
  4. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae "48.3 Yunnan, People's Republic of China; 2. Dark Red Beds of the Lower Lufeng Series," in Weishampel, et al. (2004). Page 534.
  5. ↑ Young, C.-C. (1982). On a primitive crocodile from Lufeng, Yunnnan. Selected Works on Yang Zhungijang. Science Press, Beijing pp. 26–28.
  6. ↑ Young, C.C., 1951, "The Lufeng saurischian fauna in China", Palaeontologia Sinica, New Series C, 13: 1-96
  7. ↑ Script error: No such module "Citation/CS1".
  8. ↑ Script error: No such module "Citation/CS1".
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  11. ↑ a b "Table 15.1," in Weishampel, et al. (2004). Page 336.
  12. ↑ a b c d Script error: No such module "Citation/CS1".
  13. ↑ a b c d e f g h i j k l m n o p q r s "48.3 Yunnan, People's Republic of China; 1. Dull Purplish Beds of the Lower Lufeng Series and 2. Dark Red Beds of the Lower Lufeng Series" in Weishampel, et al. (2004). Page 534.
  14. ↑ a b c d e f g h i j k l m n o "48.3 Yunnan, People's Republic of China; 1. Dull Purplish Beds of the Lower Lufeng Series," in Weishampel, et al. (2004). Page 534.
  15. ↑ a b "Table 12.1," in Weishampel, et al. (2004). Page 234.
  16. ↑ a b "Table 12.1," in Weishampel, et al. (2004). Page 236.
  17. ↑ a b c d e Script error: No such module "Citation/CS1".
  18. ↑ Script error: No such module "Citation/CS1".
  19. ↑ Script error: No such module "Citation/CS1".
  20. ↑ "Table 7.1," in Weishampel, et al. (2004). Page 152.
  21. ↑ Script error: No such module "Citation/CS1".
  22. ↑ M. N. Bien. 1940. Discovery of Triassic saurischian and primitive mammalian remains at Lufeng, Yunnan. Bulletin of the Geological Society of China 20(3/4):225-234
  23. ↑ Script error: No such module "Citation/CS1".
  24. ↑ a b c d e Script error: No such module "Citation/CS1".
  25. ↑ a b c Script error: No such module "Citation/CS1".
  26. ↑ Script error: No such module "Citation/CS1".
  27. ↑ a b c Script error: No such module "Citation/CS1".
  28. ↑ Script error: No such module "Citation/CS1".

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Bibliography

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  • Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. Template:ISBN.
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