Coffea: Difference between revisions

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|image_caption = Flowering branches of ''[[Coffea arabica]]''
|image_caption = Flowering branches of ''[[Coffea arabica]]''
|taxon = Coffea
|taxon = Coffea
|authority = [[Carl Linnaeus|L.]]
|authority = [[Carl Linnaeus|L.]]<ref name="POWO">{{cite web |url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:325985-2 |title=''Coffea'' L. |website=Plants of the World Online |publisher=Board of Trustees of the Royal Botanic Gardens, Kew |year=2026 |access-date=13 January 2026}}</ref>
|type_species = ''[[Coffea arabica]]''
|type_species = ''[[Coffea arabica]]''
|type_species_authority = [[Carl Linnaeus|L.]]
|type_species_authority = [[Carl Linnaeus|L.]]
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}}
}}


'''''Coffea''''' is a [[genus]] of [[flowering plant]]s in the [[Family (biology)|family]] [[Rubiaceae]]. ''Coffea'' species are [[shrub]]s or small [[tree]]s native to tropical and southern [[Africa]] and tropical [[Asia]]. The [[seed]]s of some species, called [[coffee]] [[coffee bean|bean]]s, are used to flavor various beverages and products. The fruits, like the seeds, contain a large amount of [[caffeine]], and have a distinct sweet taste.
'''''Coffea''''' is a [[genus]] of [[flowering plant]]s in the [[Family (biology)|family]] [[Rubiaceae]]. ''Coffea'' species are shrubs or small trees native to tropical and southern Africa and tropical Asia. The seeds of some species, called coffee beans, are roasted and ground to brew into various [[coffee]] beverages. The fruits, like the seeds, contain a large amount of caffeine, and have a distinct sweet taste.


The plant ranks as one of the world's most valuable and widely traded [[commodity crop]]s and is an important [[export]] product of several countries, including those in Central and South America, the Caribbean and Africa. The coffee trade relies heavily on two of the over 120 species, ''[[Coffea arabica]]'' (commonly known simply as "Arabica"), which accounts for 60–80% of the world's coffee production, and ''[[Coffea canephora]]'' (known as "[[Robusta coffee|Robusta]]"), which accounts for about 20–40%.<ref name=":0" /><ref name="CoffeeProduction" />  
The plant ranks as one of the world's most valuable and widely traded [[commodity crop]]s and is an important export product of several countries, including those in Central and South America, the Caribbean and Africa. The coffee trade relies heavily on two of the over 120 species, ''[[Coffea arabica]]'' (commonly known simply as "Arabica"), which accounts for 60–80% of the world's coffee production, and ''[[Coffea canephora]]'' (known as "[[Robusta coffee|Robusta]]"), which accounts for about 20–40%.<ref name=":0" /><ref name="CoffeeProduction" />  


Both coffee species are vulnerable to [[Effects of climate change on agriculture|shifts, caused by climate change, in their growing zones]], which are likely to result in a decline in production in some of the most important growing regions.<ref name="laderach">{{cite journal |last1=Läderach |first1=Peter |last2=Ramirez–Villegas |first2=Julian |last3=Navarro-Racines |first3=Carlos |last4=Zelaya |first4=Carlos |last5=Martinez–Valle |first5=Armando |last6=Jarvis |first6=Andy |title=Climate change adaptation of coffee production in space and time |journal=Climatic Change |volume=141 |issue=1 |pages=47–62 |date=26 October 2016 |issn=0165-0009 |doi=10.1007/s10584-016-1788-9 |doi-access=free|hdl=10568/77563 |hdl-access=free }}</ref><ref name="Moat">{{cite journal |last1=Moat |first1=Justin |last2=Williams |first2=Jenny |last3=Baena |first3=Susana |last4=Wilkinson |first4=Timothy |last5=Gole |first5=Tadesse W. |last6=Challa |first6=Zeleke K. |last7=Demissew |first7=Sebsebe |last8=Davis |first8=Aaron P. |title=Resilience potential of the Ethiopian coffee sector under climate change |url=https://www.researchgate.net/publication/317894875 |journal=Nature Plants |volume=3 |issue=7 |pages=17081 |date=19 June 2017 |issn=2055-0278 |pmid=28628132 |s2cid=6873955 |doi=10.1038/nplants.2017.81|bibcode=2017NatPl...317081M }}</ref><ref name="worland">{{cite magazine |last=Worland |first=Justin |date=21 June 2018 |title=Your Morning Cup of Coffee Is in Danger. Can the Industry Adapt in Time? |url=https://time.com/5318245/coffee-industry-climate-change/ |url-status=live |magazine=[[Time (magazine)|Time]] |archive-url=https://web.archive.org/web/20190804074707/https://time.com/5318245/coffee-industry-climate-change/ |archive-date=4 August 2019 |access-date=13 October 2019}}</ref><ref>{{cite book |last1=Lashermes |first1=Philippe |last2=Andrade |first2=Alan Carvalho |last3=Etienne |first3=Hervé |title=Genomics of Tropical Crop Plants |date=2008 |publisher=Springer |location=New York |isbn=978-0-387-71219-2 |pages=203–226 |doi=10.1007/978-0-387-71219-2_9 |editor1=Paul H. Moore |editor2=Ray Ming |language=en |volume=1 |chapter=Genomics of Coffee One of the World’s Largest Traded Commodities |quote=Consequently, bean quality may be lowered and supply become less stable to adversely affect the coffee market of consumer countries. Furthermore, in the light of climate changes and increasing awareness of the negative impacts of the nonsustainable use of natural resources, coffee production will have to evolve. This is particularly relevant for perennial plants such as coffee whose productive life is very long and for which rapid genetic gains are tedious to obtain.}}</ref>
Both coffee species are vulnerable to [[Effects of climate change on agriculture|shifts, caused by climate change, in their growing zones]], which are likely to result in a decline in production in some of the most important growing regions.<ref name="laderach">{{cite journal |last1=Läderach |first1=Peter |last2=Ramirez–Villegas |first2=Julian |last3=Navarro-Racines |first3=Carlos |last4=Zelaya |first4=Carlos |last5=Martinez–Valle |first5=Armando |last6=Jarvis |first6=Andy |title=Climate change adaptation of coffee production in space and time |journal=Climatic Change |volume=141 |issue=1 |pages=47–62 |date=26 October 2016 |issn=0165-0009 |doi=10.1007/s10584-016-1788-9 |doi-access=free|hdl=10568/77563 |hdl-access=free }}</ref><ref name="Moat">{{cite journal |last1=Moat |first1=Justin |last2=Williams |first2=Jenny |last3=Baena |first3=Susana |last4=Wilkinson |first4=Timothy |last5=Gole |first5=Tadesse W. |last6=Challa |first6=Zeleke K. |last7=Demissew |first7=Sebsebe |last8=Davis |first8=Aaron P. |title=Resilience potential of the Ethiopian coffee sector under climate change |url=https://www.researchgate.net/publication/317894875 |journal=Nature Plants |volume=3 |issue=7 |pages=17081 |date=19 June 2017 |issn=2055-0278 |pmid=28628132 |s2cid=6873955 |doi=10.1038/nplants.2017.81|bibcode=2017NatPl...317081M }}</ref><ref name="worland">{{cite magazine |last=Worland |first=Justin |date=21 June 2018 |title=Your Morning Cup of Coffee Is in Danger. Can the Industry Adapt in Time? |url=https://time.com/5318245/coffee-industry-climate-change/ |url-status=live |magazine=[[Time (magazine)|Time]] |archive-url=https://web.archive.org/web/20190804074707/https://time.com/5318245/coffee-industry-climate-change/ |archive-date=4 August 2019 |access-date=13 October 2019}}</ref><ref>{{cite book |last1=Lashermes |first1=Philippe |last2=Andrade |first2=Alan Carvalho |last3=Etienne |first3=Hervé |title=Genomics of Tropical Crop Plants |date=2008 |publisher=Springer |location=New York |isbn=978-0-387-71219-2 |pages=203–226 |doi=10.1007/978-0-387-71219-2_9 |editor1=Paul H. Moore |editor2=Ray Ming |language=en |volume=1 |chapter=Genomics of Coffee One of the World’s Largest Traded Commodities |quote=Consequently, bean quality may be lowered and supply become less stable to adversely affect the coffee market of consumer countries. Furthermore, in the light of climate changes and increasing awareness of the negative impacts of the nonsustainable use of natural resources, coffee production will have to evolve. This is particularly relevant for perennial plants such as coffee whose productive life is very long and for which rapid genetic gains are tedious to obtain.}}</ref>
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=== Species ===
=== Species ===
As of May 2024, [[Plants of the World Online]] includes:<ref name=POWO>[https://powo.science.kew.org/taxon/urn%3Alsid%3Aipni.org%3Anames%3A325985-2 Plants of the World Online: ''Coffea'' L.] (retrieved 31 May 2024)</ref>
{{As of|2026|01}}, [[Plants of the World Online]] accepts the following 133 species and one hybrid:{{R|POWO}}
{{div col|colwidth=20em}}
 
# ''[[Coffea abbayesii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
{{columns-list|colwidth=22em|
# ''[[Coffea affinis]]'' <small>[[Émile Auguste Joseph De Wildeman|De Wild.]]</small>
*''[[Coffea abbayesii]]'' {{small|J.-F.Leroy}}
# ''[[Coffea alleizettii]]'' <small>Dubard</small>
*''[[Coffea affinis]]'' {{small|De Wild.}}
# ''[[Coffea ambanjensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea alleizettii]]'' {{small|Dubard}}
# ''[[Coffea ambongenis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]] ex A. P. Davis</small>
*''[[Coffea ambanjensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea andrambovatensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea ambongensis]]'' {{small|J.-F.Leroy ex A.P.Davis & Rakotonas.}}
# ''[[Coffea ankaranensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]] ex A. P. Davis</small>
*''[[Coffea andrambovatensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea anthonyi]]'' <small>Stoff. & F. Anthony</small>
*''[[Coffea ankaranensis]]'' {{small|J.-F.Leroy ex A.P.Davis & Rakotonas.}}
# ''[[Coffea arabica]]'' <small>[[Carl Linnaeus|L.]]</small>
*''[[Coffea anthonyi]]'' {{small|Stoff. & F.Anthony}}
# ''[[Coffea arenesiana]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea arabica]]'' {{small|L.}}
# ''[[Coffea augagneurii]]'' <small>Dubard</small>
*''[[Coffea arenesiana]]'' {{small|J.-F.Leroy}}
# ''[[Coffea bakossii]]'' <small>Cheek & [[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea augagneurii]]'' {{small|Dubard}}
# ''[[Coffea benghalensis]]'' <small>B. Heyne ex [[Josef August Schultes|Schult.]]</small>
*''[[Coffea bakossii]]'' {{small|Cheek & Bridson}}
# ''[[Coffea bertrandii]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea benghalensis]]'' {{small|B.Heyne ex Roth}}
# ''[[Coffea betamponensis]]'' <small>Portères & [[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea bertrandii]]'' {{small|A.Chev.}}
# ''[[Coffea bissetiae]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea betamponensis]]'' {{small|Portères & J.-F.Leroy}}
# ''[[Coffea boinensis]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea bissetiae]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea boiviniana]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea boinensis]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea bonnieri]]'' <small>Dubard</small>
*''[[Coffea boiviniana]]'' {{small|(Baill.) Drake}}
# ''[[Coffea brassii]]'' <small>([[Jean-François Leroy (botanist)|J.-F. Leroy]]) A. P. Davis</small>
*''[[Coffea bonnieri]]'' {{small|Dubard}}
# ''[[Coffea brevipes]]'' <small>[[William Philip Hiern|Hiern]]</small>
*''[[Coffea brassii]]'' {{small|(J.-F.Leroy) A.P.Davis}}
# ''[[Coffea bridsoniae]]'' <small>A. P. Davis & Mvungi</small>
*''[[Coffea brevipes]]'' {{small|Hiern}}
# ''[[Coffea buxifolia]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea bridsoniae]]'' {{small|A.P.Davis & Mvungi}}
# ''[[Coffea callmanderi]]'' {{Au|A. P. Davis & Rakotonas.}}
*''[[Coffea buxifolia]]'' {{small|A.Chev.}}
# ''[[Coffea canephora]]'' (''"Coffea robusta"'') <small>Pierre ex A. Froehner</small>
*''[[Coffea callmanderi]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea carrissoi]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea canephora]]'' {{small|Pierre ex A.Froehner}}
# ''[[Coffea charrieriana]]'' <small>Stoff. & F. Anthony</small>
*''[[Coffea carrissoi]]'' {{small|A.Chev.}}
# ''[[Coffea cochinchinensis]]'' <small>Pierre ex Pit.</small>
*''[[Coffea charrieriana]]'' {{small|Stoff. & F.Anthony}}
# ''[[Coffea commersoniana]]'' <small>([[Henri Ernest Baillon|Baill.]]) [[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea cochinchinensis]]'' {{small|Pierre ex Pit.}}
# ''[[Coffea congensis]]'' <small>A. Froehner</small>
*''[[Coffea commersoniana]]'' {{small|(Baill.) A.Chev.}}
# ''[[Coffea costatifructa]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea congensis]]'' {{small|A.Froehner}}
# ''[[Coffea coursiana]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea costatifructa]]'' {{small|Bridson}}
# ''[[Coffea dactylifera]]'' <small>[[Elmar Robbrecht|Robbr.]] & Stoff.</small>
*''[[Coffea coursiana]]'' {{small|J.-F.Leroy}}
# ''[[Coffea darainensis]]'' {{Au|A. P. Davis & Rakotonas.}}
*''[[Coffea dactylifera]]'' {{small|Robbr. & Stoff.}}
# ''[[Coffea decaryana]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea darainensis]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea dubardii]]'' <small>Jum.</small>
*''[[Coffea decaryana]]'' {{small|J.-F.Leroy}}
# ''[[Coffea ebracteolata]]'' <small>([[William Philip Hiern|Hiern]]) Brenan</small>
*''[[Coffea dewevrei]]'' {{small|De Wild. & T.Durand}}
# ''[[Coffea eugenioides]]'' <small>[[Spencer Le Marchant Moore|S. Moore]]</small>
*''[[Coffea dubardii]]'' {{small|Jum.}}
# ''[[Coffea fadenii]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea ebracteolata]]'' {{small|(Hiern) Brenan}}
# ''[[Coffea farafanganensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea eugenioides]]'' {{small|S.Moore}}
# ''[[Coffea floresiana]]'' <small>Boerl.</small>
*''[[Coffea fadenii]]'' {{small|Bridson}}
# ''[[Coffea fotsoana]]'' <small>Stoff. & Sonké</small>
*''[[Coffea farafanganensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea fragilis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea floresiana]]'' {{small|Boerl.}}
# ''[[Coffea fragrans]]'' <small>[[Nathaniel Wallich|Wall.]] ex [[Joseph Dalton Hooker|Hook. f.]]</small>
*''[[Coffea fotsoana]]'' {{small|Stoff. & Sonké}}
# ''[[Coffea gallienii]]'' <small>Dubard</small>
*''[[Coffea fragilis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea grevei]]'' <small>[[Emmanuel Drake del Castillo|Drake]] ex [[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea fragrans]]'' {{small|Wall. ex Hook.f.}}
# ''[[Coffea heimii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea gallienii]]'' {{small|Dubard}}
# ''[[Coffea × heterocalyx]]'' {{Au|Stoff.}}
*''[[Coffea grevei]]'' {{small|Drake ex A.Chev.}}
# ''[[Coffea homollei]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea heimii]]'' {{small|J.-F.Leroy}}
# ''[[Coffea horsfieldiana]]'' <small>[[Friedrich Anton Wilhelm Miquel|Miq.]]</small>
[[Coffea × heterocalyx|''Coffea'' × ''heterocalyx'']] {{small|Stoff.}}
# ''[[Coffea humbertii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea homollei]]'' {{small|J.-F.Leroy}}
# ''[[Coffea humblotiana]]'' <small>[[Henri Ernest Baillon|Baill.]]</small>
*''[[Coffea horsfieldiana]]'' {{small|Miq.}}
# ''[[Coffea humilis]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea humbertii]]'' {{small|J.-F.Leroy}}
# ''[[Coffea jumellei]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea humblotiana]]'' {{small|Baill.}}
# ''[[Coffea kalobinonensis]]'' {{Au|A. P. Davis & Rakotonas.}}
*''[[Coffea humilis]]'' {{small|A.Chev.}}
# ''[[Coffea kapakata]]'' <small>([[Auguste Jean Baptiste Chevalier|A. Chev.]]) [[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea jumellei]]'' {{small|J.-F.Leroy}}
# ''[[Coffea kianjavatensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea kalobinonensis]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea kihansiensis]]'' <small>A. P. Davis & Mvungi</small>
*''[[Coffea kapakata]]'' {{small|(A.Chev.) Bridson}}
# ''[[Coffea kimbozensis]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea kianjavatensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea kivuensis]]'' <small>Lebrun</small>
*''[[Coffea kihansiensis]]'' {{small|A.P.Davis & Mvungi}}
# ''[[Coffea labatii]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea kimbozensis]]'' {{small|Bridson}}
# ''[[Coffea lancifolia]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea kivuensis]]'' {{small|Lebrun}}
# ''[[Coffea lebruniana]]'' <small>Germ. & Kester</small>
*''[[Coffea klainei]]'' {{small|Pierre ex De Wild.}}
# ''[[Coffea leonimontana]]'' <small>Stoff.</small>
*''[[Coffea labatii]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea leroyi]]'' <small>A. P. Davis</small>
*''[[Coffea lancifolia]]'' {{small|A.Chev.}}
# ''[[Coffea liaudii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]] ex A. P. Davis</small>
*''[[Coffea lebruniana]]'' {{small|Germ. & Kesler}}
# ''[[Coffea liberica]]'' <small>[[William Philip Hiern|Hiern]]</small>
*''[[Coffea leonimontana]]'' {{small|Stoff.}}
# ''[[Coffea ligustroides]]'' <small>[[Spencer Le Marchant Moore|S. Moore]]</small>
*''[[Coffea leroyi]]'' {{small|A.P.Davis}}
# ''[[Coffea littoralis]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea liaudii]]'' {{small|J.-F.Leroy ex A.P.Davis}}
# ''[[Coffea lulandoensis]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea liberica]]'' {{small|W.Bull}}
# ''[[Coffea mabesae]]'' <small>(Elmer) [[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea ligustroides]]'' {{small|S.Moore}}
# ''[[Coffea macrocarpa]]'' <small>[[Achille Richard|A. Rich.]]</small>
*''[[Coffea littoralis]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea madurensis]]'' <small>[[Johannes Elias Teijsmann|Teijsm.]] & [[Simon Binnendijk|Binn.]] ex Koord.</small>
*''[[Coffea lulandoensis]]'' {{small|Bridson}}
# ''[[Coffea magnistipula]]'' <small>Stoff. & [[Elmar Robbrecht|Robbr.]]</small>
*''[[Coffea mabesae]]'' {{small|(Elmer) J.-F.Leroy}}
# ''[[Coffea malabarica]]'' <small>(Sivar., Biju & P. Mathew) A.P.Davis</small>
*''[[Coffea macrocarpa]]'' {{small|A.Rich.}}
# ''[[Coffea mangoroensis]]'' <small>Portères</small>
*''[[Coffea madurensis]]'' {{small|Teijsm. & Binn. ex Koord.}}
# ''[[Coffea mannii]]'' <small>([[Joseph Dalton Hooker|Hook. f.]]) A. P. Davis</small>
*''[[Coffea magnistipula]]'' {{small|Stoff. & Robbr.}}
# ''[[Coffea manombensis]]'' <small>A. P. Davis</small>
*''[[Coffea malabarica]]'' {{small|(Sivar., Biju & P.Mathew) A.P.Davis}}
# ''[[Coffea mapiana]]'' <small>Sonké, Nguembou & A P. Davis</small>
*''[[Coffea mangoroensis]]'' {{small|Portères}}
# ''[[Coffea mauritiana]]'' <small>[[Jean-Baptiste Lamarck|Lam.]]</small>
*''[[Coffea mannii]]'' {{small|(Hook.f.) A.P.Davis}}
# ''[[Coffea mayombensis]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea manombensis]]'' {{small|A.P.Davis}}
# ''[[Coffea mcphersonii]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea mapiana]]'' {{small|Sonké, Nguembou & A.P.Davis}}
# ''[[Coffea melanocarpa]]'' <small>[[Friedrich Welwitsch|Welw.]] ex [[William Philip Hiern|Hiern]]</small>
*''[[Coffea mauritiana]]'' {{small|Lam.}}
# ''[[Coffea merguensis]]'' <small>[[Henry Nicholas Ridley|Ridl.]]</small>
*''[[Coffea mayombensis]]'' {{small|A.Chev.}}
# ''[[Coffea microdubardii]]'' {{Au|A. P. Davis & Rakotonas.}}
*''[[Coffea mcphersonii]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea millotii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea melanocarpa]]'' {{small|Welw. ex Hiern}}
# ''[[Coffea minutiflora]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea merguensis]]'' {{small|Ridl.}}
# ''[[Coffea mogenetii]]'' <small>Dubard</small>
*''[[Coffea microdubardii]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea mongensis]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea millotii]]'' {{small|J.-F.Leroy}}
# ''[[Coffea montekupensis]]'' <small>Stoff.</small>
*''[[Coffea minutiflora]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea montis-sacri]]'' <small>A. P. Davis</small>
*''[[Coffea mogenetii]]'' {{small|Dubard}}
# ''[[Coffea moratii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]] ex A. P. Davis & Rakotonas.</small>
*''[[Coffea mongensis]]'' {{small|Bridson}}
# ''[[Coffea mufindiensis]]'' <small>[[John Hutchinson (botanist)|Hutch.]] ex [[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea montekupensis]]'' {{small|Stoff.}}
# ''[[Coffea myrtifolia]]'' <small>([[Achille Richard|A.Rich.]] ex [[Augustin Pyramus de Candolle|DC.]]) [[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea montis-sacri]]'' {{small|A.P.Davis}}
# ''[[Coffea namorokensis]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea moratii]]'' {{small|J.-F.Leroy ex A.P.Davis & Rakotonas.}}
# ''[[Coffea neobridsoniae]]'' <small>A. P. Davis</small>
*''[[Coffea mufindiensis]]'' {{small|Hutch. ex Bridson}}
# ''[[Coffea neoleroyi]]'' <small>A. P. Davis</small>
*''[[Coffea myrtifolia]]'' {{small|(A.Rich. ex DC.) J.-F.Leroy}}
# ''[[Coffea perrieri]]'' <small>[[Emmanuel Drake del Castillo|Drake]] ex Jum. & H. Perrier</small>
*''[[Coffea namorokensis]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea pervilleana]]'' <small>([[Henri Ernest Baillon|Baill.]]) [[Emmanuel Drake del Castillo|Drake]]</small>
*''[[Coffea neobridsoniae]]'' {{small|A.P.Davis}}
# ''[[Coffea pocsii]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea neoleroyi]]'' {{small|A.P.Davis}}
# ''[[Coffea pseudozanguebariae]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea perrieri]]'' {{small|Drake ex Jum. & H.Perrier}}
# ''[[Coffea pterocarpa]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea pervilleana]]'' {{small|(Baill.) Drake}}
# ''[[Coffea pustulata]]'' {{Au|A. P. Davis & Rakotonas.}}
*''[[Coffea pocsii]]'' {{small|Bridson}}
# ''[[Coffea racemosa]]'' <small>[[João de Loureiro|Lour.]]</small>
*''[[Coffea pseudozanguebariae]]'' {{small|Bridson}}
# ''[[Coffea rakotonasoloi]]'' <small>A. P. Davis</small>
*''[[Coffea pterocarpa]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea ratsimamangae]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]] ex A. P. Davis & Rakotonas.</small>
*''[[Coffea pustulata]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea resinosa]]'' <small>([[Joseph Dalton Hooker|Hook. f.]]) Radlk.</small>
*''[[Coffea racemosa]]'' {{small|Lour.}}
# ''[[Coffea rhamnifolia]]'' <small>([[Emilio Chiovenda|Chiov.]]) [[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea rakotonasoloi]]'' {{small|A.P.Davis}}
# ''[[Coffea richardii]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea ratsimamangae]]'' {{small|J.-F.Leroy ex A.P.Davis & Rakotonas.}}
# ''[[Coffea rizetiana]]'' {{Au|Stoff. & Noirot}}
*''[[Coffea resinosa]]'' {{small|(Hook.f.) Radlk.}}
# ''[[Coffea rupicola]]'' {{Au|A. P. Davis & Rakotonas.}}
*''[[Coffea rhamnifolia]]'' {{small|(Chiov.) Bridson}}
# ''[[Coffea sahafaryensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea richardii]]'' {{small|J.-F.Leroy}}
# ''[[Coffea sakarahae]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea rizetiana]]'' {{small|Stoff. & Noirot}}
# ''[[Coffea salvatrix]]'' <small>Swynn. & Philipson</small>
*''[[Coffea rupicola]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea sambavensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]] ex A. P Davis & Rakotonas.</small>
*''[[Coffea sahafaryensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea sapinii]]'' <small>([[Émile Auguste Joseph De Wildeman|De Wild.]]) A. P. Davis</small>
*''[[Coffea sakarahae]]'' {{small|J.-F.Leroy}}
# ''[[Coffea schliebenii]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea salvatrix]]'' {{small|Swynn. & Philipson}}
# ''[[Coffea semsei]]'' <small>([[Diane Mary Bridson|Bridson]]) A. P. Davis</small>
*''[[Coffea sambavensis]]'' {{small|J.-F.Leroy ex A.P.Davis & Rakotonas.}}
# ''[[Coffea sessiliflora]]'' <small>[[Diane Mary Bridson|Bridson]]</small>
*''[[Coffea sapinii]]'' {{small|(De Wild.) A.P.Davis}}
# ''[[Coffea stenophylla]]'' <small>[[George Don|G. Don]]</small>
*''[[Coffea schliebenii]]'' {{small|Bridson}}
# ''[[Coffea tetragona]]'' <small>Jum. & H. Perrier</small>
*''[[Coffea semsei]]'' {{small|(Bridson) A.P.Davis}}
# ''[[Coffea togoensis]]'' <small>[[Auguste Jean Baptiste Chevalier|A. Chev.]]</small>
*''[[Coffea sessiliflora]]'' {{small|Bridson}}
# ''[[Coffea toshii]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea stenophylla]]'' {{small|G.Don}}
# ''[[Coffea travancorensis]]'' <small>[[Robert Wight|Wight]] & [[George Arnott Walker-Arnott|Arn.]]</small>
*''[[Coffea tetragona]]'' {{small|Jum. & H.Perrier}}
# ''[[Coffea tricalysioides]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea togoensis]]'' {{small|A.Chev.}}
# ''[[Coffea tsirananae]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea toshii]]'' {{small|A.P.Davis & Rakotonas.}}
# ''[[Coffea vatovavyensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea travancorensis]]'' {{small|Wight & Arn.}}
# ''[[Coffea vavateninensis]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea tricalysioides]]'' {{small|J.-F.Leroy}}
# ''[[Coffea vianneyi]]'' <small>[[Jean-François Leroy (botanist)|J.-F. Leroy]]</small>
*''[[Coffea tsirananae]]'' {{small|J.-F.Leroy}}
# ''[[Coffea vohemarensis]]'' <small>A. P. Davis & Rakotonas.</small>
*''[[Coffea vatovavyensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea wightiana]]'' <small>[[Nathaniel Wallich|Wall.]] ex [[Robert Wight|Wight]] & [[George Arnott Walker-Arnott|Arn.]]</small>
*''[[Coffea vavateninensis]]'' {{small|J.-F.Leroy}}
# ''[[Coffea zanguebariae]]'' <small>[[João de Loureiro|Lour.]]</small>
*''[[Coffea vianneyi]]'' {{small|J.-F.Leroy}}
{{div col end}}
*''[[Coffea vohemarensis]]'' {{small|A.P.Davis & Rakotonas.}}
*''[[Coffea wightiana]]'' {{small|Wall. ex Wight & Arn.}}
*''[[Coffea zanguebariae]]'' {{small|Lour.}}
}}


== Ecology ==
== Ecology ==

Latest revision as of 21:15, 12 March 2026

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Coffea is a genus of flowering plants in the family Rubiaceae. Coffea species are shrubs or small trees native to tropical and southern Africa and tropical Asia. The seeds of some species, called coffee beans, are roasted and ground to brew into various coffee beverages. The fruits, like the seeds, contain a large amount of caffeine, and have a distinct sweet taste.

The plant ranks as one of the world's most valuable and widely traded commodity crops and is an important export product of several countries, including those in Central and South America, the Caribbean and Africa. The coffee trade relies heavily on two of the over 120 species, Coffea arabica (commonly known simply as "Arabica"), which accounts for 60–80% of the world's coffee production, and Coffea canephora (known as "Robusta"), which accounts for about 20–40%.[1][2]

Both coffee species are vulnerable to shifts, caused by climate change, in their growing zones, which are likely to result in a decline in production in some of the most important growing regions.[3][4][5][6]

Taxonomy

The genus Coffea and the type species Coffea arabica were first described by Carl Linnaeus in 1753.[7] New species of Coffea are still being identified in the 2000s. In 2008 and 2009, researchers from the Royal Botanic Gardens, Kew, named seven from the mountains of northern Madagascar, including C. ambongensis, C. boinensis, C. labatii, C. pterocarpa, C. bissetiae, and C. namorokensis.[8]

In 2008, two new species were discovered in Cameroon: Coffea charrieriana, which is caffeine-free, and Coffea anthonyi.[9] By crossing the new species with other known coffees, two new features might be introduced to cultivated coffee plants: beans without caffeine and self-pollination.

In 2011, Coffea absorbed the twenty species of the former genus Psilanthus due to the morphological and genetic similarities between the two genera.[10] Historically, the two have been considered distinct genera due to differences in the length of the corolla tube and the anther arrangement: Coffea with a short corolla tube and exserted style and anthers; Psilanthus with a long corolla tube and included anthers. However, these characteristics were not present in all species of either respective genus, making the two genera overwhelmingly similar in both morphology and genetic sequence. This transfer expanded Coffea from 104 species to 124, and extended its native distribution to tropical Asia and Australasia.

The coffee genome was published in 2014, with more than 25,000 genes identified. This revealed that coffee plants make caffeine using a different set of genes from those found in tea, cacao and other such plants.[11]

A robust and almost fully resolved phylogeny of the entire genus was published in 2017.[12] In addition to resolving the relationships of Coffea species, this study's results suggest Africa or Asia as the likely ancestral origin of Coffea and point to several independent radiations across Africa, Asia, and the Western Indian Ocean Islands.

In 2020, a technique of DNA fingerprinting, or genetic authentication of plant material, was proven effective for coffee.[13] For the study, scientists used DNA extraction and SSR marker analysis. This technique or similar ones may allow for several improvements to coffee production such as improved information for farmers as to the susceptibility of their coffee plants to pests and disease, a professionalized coffee seed system, and transparency and traceability for buyers of green, un-roasted coffee.

Species

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Ecology

The caffeine in coffee beans serves as a toxic substance that protects against insects and other pests, a form of natural plant defense against herbivory. Caffeine simultaneously attracts pollinators, specifically honeybees, by creating an olfactory memory that signals bees to return to the plant's flowers.[12] Not all Coffea species contain caffeine, and the earliest species had little or no caffeine content. Caffeine has evolved independently in multiple lineages of Coffea in Africa, perhaps in response to high pest predation in the humid environments of West-Central Africa.[12]

Caffeine has also evolved independently in the more distantly related genera Theobroma (cacao) and Camellia (tea).[14] This suggests that caffeine production is an adaptive trait in coffee and plant evolution. The fruit and leaves also contain caffeine, and can be used to make coffee cherry tea and coffee-leaf tea. The fruit is also used in many brands of soft drink as well as pre-packaged teas.[15][16][17][18]

Several insect pests affect coffee production, including the coffee borer beetle (Hypothenemus hampei) and the coffee leafminer (Leucoptera caffeina).

Coffee is used as a food plant by the larvae of some Lepidoptera (butterfly and moth) species, Dalcera abrasa, turnip moth and some members of the genus Endoclita, including E. damor and E. malabaricus.

Cultivation and use

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Freshly harvested coffee cherries
Freshly harvested coffee cherries
File:CoffeeBerry.jpg
Coffea fruits, Bali

There are over 130 species of Coffea, which is grown from seed. The two most popular are Coffea arabica (commonly known simply as "Arabica"), which accounts for 60–80% of the world's coffee production, and Coffea canephora (known as "Robusta"), which accounts for about 20–40%.[1][2] C. arabica is preferred for its sweeter taste, while C. canephora has a higher caffeine content. C. arabica has its origins in the highlands of Ethiopia and the Boma Plateau of Sudan, and came about as the result of a hybrid between C. canephora and C. eugenioides.[19]

The trees produce edible red or purple fruits that are either epigynous berries or indehiscent drupes.[20] The fruit is often referred to as a "coffee cherry", and it contains two seeds, called "coffee beans".[21] Despite these terms, coffee is neither a true cherry (the fruit of certain species in the genus Prunus) nor a true bean (seeds from plants in the family Fabaceae).

In any coffee crop, about 5–10% of fruits contain only a single bean. Called a peaberry, it is smaller and rounder than a normal coffee bean.

When grown in the tropics, coffee is a vigorous bush or small tree that usually grows to a height of Script error: No such module "convert".. Most commonly cultivated coffee species grow best at high elevations, but do not tolerate freezing temperatures.[22]

The Coffea arabica tree grows fruit after three to five years, producing for an average of 50 to 60 years, though up to 100 years is possible.[23] The white flowers are highly scented. The fruit takes about nine months to ripen.

File:Coffe flower.jpg
Coffea flower
File:Pollen grains of Coffee plant.jpg
Pollen grains of Coffee plant
File:Coffee cherry cross section.JPG
Coffea fruit cross section
File:Coffea racemosa00.jpg
Coffea racemosa fruits
File:Coffee germination.jpg
Coffea arabica beans germinating
File:Coffee arabica flower.jpg
Coffea arabica flowers
File:Coffee arabica cherry.jpg
Ripe Coffea arabica fruits
File:Open coffee cherry.jpg
Beans inside a Coffea arabica fruit
File:咖啡樹-草坪頭 Coffea 20220203134728.jpg
Coffea branches

References

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External links

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