Clupeiformes: Difference between revisions

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The former order of '''Isospondyli''' was subsumed mostly by Clupeiformes,<ref>{{cite book|title=Journal of Ichthyology|url=https://books.google.com/books?id=FhAkAQAAIAAJ|volume=46|year=2006|publisher=Scripta|page=S40|quote= within Isospondyli (= Clupeiformes ''[[sensu lato|s. lato]]'')}}</ref> but some isospondylous fishes (isospondyls) were assigned to [[Osteoglossiformes]], [[Salmoniformes]], [[Cetomimiformes]], etc.<ref>{{cite book|author=lfonso L. Rojo|title=Dictionary of Evolutionary Fish Osteology|url=https://books.google.com/books?id=Uc03DwAAQBAJ&pg=PT170|year=2017|publisher=CRC |isbn=978-1-351-36604-5|page=170|quote= Under the name '''Isospondyli''', Regan (1909) grouped the fishes having the verterbrae immediately after the skull similar in shape to the remaining ones, in contrast to the [[ostariophysan]]s, in which the anterior vertebrae are greatly modified. Modern classifications have rejected this artificially constructed group, and the fishes previously assigned to it have been distributed among different orders (Clupeiformes, Osteoglossiformes, Salmoniformes, Cetomimiformes, etc.)}}</ref>
The former order of '''Isospondyli''' was subsumed mostly by Clupeiformes,<ref>{{cite book|title=Journal of Ichthyology|url=https://books.google.com/books?id=FhAkAQAAIAAJ|volume=46|year=2006|publisher=Scripta|page=S40|quote= within Isospondyli (= Clupeiformes ''[[sensu lato|s. lato]]'')}}</ref> but some isospondylous fishes (isospondyls) were assigned to [[Osteoglossiformes]], [[Salmoniformes]], [[Cetomimiformes]], etc.<ref>{{cite book|author=lfonso L. Rojo|title=Dictionary of Evolutionary Fish Osteology|url=https://books.google.com/books?id=Uc03DwAAQBAJ&pg=PT170|year=2017|publisher=CRC |isbn=978-1-351-36604-5|page=170|quote= Under the name '''Isospondyli''', Regan (1909) grouped the fishes having the verterbrae immediately after the skull similar in shape to the remaining ones, in contrast to the [[ostariophysan]]s, in which the anterior vertebrae are greatly modified. Modern classifications have rejected this artificially constructed group, and the fishes previously assigned to it have been distributed among different orders (Clupeiformes, Osteoglossiformes, Salmoniformes, Cetomimiformes, etc.)}}</ref>


Their [[sister group]] were the extinct [[Ellimmichthyiformes]], which were dominant throughout much of the [[Cretaceous]] and into the [[Paleogene]],<ref name=":02">{{Cite book |last1=Nelson |first1=Joseph S. |url=http://dx.doi.org/10.1002/9781119174844 |title=Fishes of the World |last2=Grande |first2=Terry C. |last3=Wilson |first3=Mark V. H. |date=2016-02-22 |publisher=Wiley |isbn=978-1-118-34233-6 |doi=10.1002/9781119174844}}</ref> and often coexisted with clupeiforms at many known localities. Both groups closely resembled each other morphologically, although the ellimmichthyiformes evolved some highly divergent body plans later in the Cretaceous.
Their [[sister group]] were the extinct [[Ellimmichthyiformes]], which were dominant throughout much of the [[Cretaceous]] and into the [[Paleogene]],<ref name=":02">{{Cite book |last1=Nelson |first1=Joseph S. |title=Fishes of the World |last2=Grande |first2=Terry C. |last3=Wilson |first3=Mark V. H. |date=2016-02-22 |publisher=Wiley |isbn=978-1-118-34233-6 |doi=10.1002/9781119174844}}</ref> and often coexisted with clupeiforms at many known localities. Both groups closely resembled each other morphologically, although the ellimmichthyiformes evolved some highly divergent body plans later in the Cretaceous.


Several fossil clupeiforms are known from the [[Early Cretaceous]] of [[South America]] that appear to be more closely allied with Clupeioidei over the [[Denticle herring|Denticipitidae]]. This suggests a very deep divergence within the [[crown group]] Clupeiformes that must have occurred during the Early Cretaceous or before.<ref name="De Figueiredo">{{cite journal |author=Francisco J. De Figueiredo |year=2009 |title=A new clupeiform fish from the Lower Cretaceous (Barremian) of Sergipe-Alagoas Basin, Northeastern Brazil |journal=[[Journal of Vertebrate Paleontology]] |volume=29 |issue=4 |pages=993–1005 |bibcode=2009JVPal..29..993D |doi=10.1671/039.029.0402 |s2cid=220436023}}</ref><ref name=":0">{{Cite journal |last1=Malabarba |first1=Maria C. |last2=Dario |first2=Fabio Di |date=2017 |title=A new predatory herring-like fish (Teleostei: Clupeiformes) from the early Cretaceous of Brazil, and implications for relationships in the Clupeoidei |url=https://academic.oup.com/zoolinnean/article/180/1/175/3799580 |journal=Zoological Journal of the Linnean Society |volume=180 |issue=1 |pages=175–194}}</ref>
Several fossil clupeiforms are known from the [[Early Cretaceous]] of [[South America]] that appear to be more closely allied with Clupeioidei over the [[Denticle herring|Denticipitidae]]. This suggests a very deep divergence within the [[crown group]] Clupeiformes that must have occurred during the Early Cretaceous or before.<ref name="De Figueiredo">{{cite journal |author=Francisco J. De Figueiredo |year=2009 |title=A new clupeiform fish from the Lower Cretaceous (Barremian) of Sergipe-Alagoas Basin, Northeastern Brazil |journal=[[Journal of Vertebrate Paleontology]] |volume=29 |issue=4 |pages=993–1005 |bibcode=2009JVPal..29..993D |doi=10.1671/039.029.0402 |s2cid=220436023}}</ref><ref name=":0">{{Cite journal |last1=Malabarba |first1=Maria C. |last2=Dario |first2=Fabio Di |date=2017 |title=A new predatory herring-like fish (Teleostei: Clupeiformes) from the early Cretaceous of Brazil, and implications for relationships in the Clupeoidei |url=https://academic.oup.com/zoolinnean/article/180/1/175/3799580 |journal=Zoological Journal of the Linnean Society |volume=180 |issue=1 |pages=175–194}}</ref>
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The order includes about 405 species in ten families:<ref name=FB>{{FishBase_order|order=Clupeiformes|year=2023|month=July}}</ref><ref>{{cite journal |author1=Lavoue ´ S |author2=Miya M |author3=Musikasinthorn P |author4=Chen W-J |author5=Nishida M |year=2013 |title=Mitogenomic Evidence for an Indo-West Pacific Origin of the Clupeoidei (Teleostei: Clupeiformes) |journal=PLOS ONE |volume=8 |issue=2 |pages=e56485 |doi=10.1371/journal.pone.0056485|pmid=23431379 |pmc=3576394|bibcode=2013PLoSO...856485L |doi-access=free }}</ref><ref>{{Catalog of Fishes |title=Eschmeyer's Catalog of Fishes Classification|url=https://www.calacademy.org/scientists/catalog-of-fishes-classification/?_gl=1*1kpq83y*_ga*NzY5Njk4MjI1LjE3MTY0NTU1NTc.*_ga_6Y72VP61VZ*MTcxNjQ1NTU1Ni4xLjAuMTcxNjQ1NTU2MS41NS4wLjA. |access-date=23 May 2024}}</ref>
The order includes about 405 species in ten families:<ref name=FB>{{FishBase_order|order=Clupeiformes|year=2023|month=July}}</ref><ref>{{cite journal |author1=Lavoue ´ S |author2=Miya M |author3=Musikasinthorn P |author4=Chen W-J |author5=Nishida M |year=2013 |title=Mitogenomic Evidence for an Indo-West Pacific Origin of the Clupeoidei (Teleostei: Clupeiformes) |journal=PLOS ONE |volume=8 |issue=2 |article-number=e56485 |doi=10.1371/journal.pone.0056485|pmid=23431379 |pmc=3576394|bibcode=2013PLoSO...856485L |doi-access=free }}</ref><ref>{{Catalog of Fishes |title=Eschmeyer's Catalog of Fishes Classification|url=https://www.calacademy.org/scientists/catalog-of-fishes-classification/?_gl=1*1kpq83y*_ga*NzY5Njk4MjI1LjE3MTY0NTU1NTc.*_ga_6Y72VP61VZ*MTcxNjQ1NTU1Ni4xLjAuMTcxNjQ1NTU2MS41NS4wLjA. |access-date=23 May 2024}}</ref>
* Order '''Clupeiformes'''  
* Order '''Clupeiformes'''  
** Genus †''[[Histiothrissa]]'' <small>[[Arthur Smith Woodward|Woodward]], 1901</small>
** Genus †''[[Histiothrissa]]'' <small>[[Arthur Smith Woodward|Woodward]], 1901</small>
Line 71: Line 71:
*** Family [[Denticipitidae]] <small>[[Herluf Stenholt Clausen|Clausen]], 1959</small> (denticle herring)
*** Family [[Denticipitidae]] <small>[[Herluf Stenholt Clausen|Clausen]], 1959</small> (denticle herring)
** Suborder [[Clupeoidei]] <small>[[Pieter Bleeker|Bleeker]], 1849</small>
** Suborder [[Clupeoidei]] <small>[[Pieter Bleeker|Bleeker]], 1849</small>
*** Genus †''[[Beksinskiella]]'' <small>[[Mateusz Granica|Granica]], [[Malgorzata Bieńowska-Wasiluki|Bieńowska-Wasiluki]] & [[Marcin Paldyna|Paldyna]], 2004</small> <ref>{{Cite journal |last1=Pałdyna |first1=Marcin |last2=Bieńkowska-Wasiluk |first2=Małgorzata |last3=Granica |first3=Mateusz |date=2024 |title=A new clupeoid genus from the Oligocene of Central Paratethys (Menilite Formation, Poland) |url=https://journals.pan.pl/dlibra/publication/148029/edition/130169 |journal=Acta Geologica Polonica |volume=74 |issue=1 |page=E5 |doi=10.24425/agp.2024.148029 |issn=0001-5709|doi-access=free }}</ref>
*** Genus †''[[Beksinskiella]]'' <small>[[Mateusz Granica|Granica]], [[Malgorzata Bieńowska-Wasiluki|Bieńowska-Wasiluki]] & [[Marcin Paldyna|Paldyna]], 2004</small><ref>{{Cite journal |last1=Pałdyna |first1=Marcin |last2=Bieńkowska-Wasiluk |first2=Małgorzata |last3=Granica |first3=Mateusz |date=2024 |title=A new clupeoid genus from the Oligocene of Central Paratethys (Menilite Formation, Poland) |url=https://journals.pan.pl/dlibra/publication/148029/edition/130169 |journal=Acta Geologica Polonica |volume=74 |issue=1 |page=E5 |doi=10.24425/agp.2024.148029 |issn=0001-5709|doi-access=free }}</ref>
*** Genus †''[[Nolfia]]'' <small>[[Francisco De Figueiredo|De Figueiredo]], 2009</small><ref>{{Cite journal |last=De Figueiredo |first=Francisco J. |date=2009-07-21 |title=A new marine clupeoid fish from the Lower Cretaceous of the Sergipe-Alagoas Basin, northeastern Brazil |url=https://biotaxa.org/Zootaxa/article/view/zootaxa.2164.1.2 |journal=Zootaxa |volume=2164 |issue=1 |pages=21–32 |doi=10.11646/zootaxa.2164.1.2 |issn=1175-5334|url-access=subscription }}</ref>
*** Genus †''[[Nolfia]]'' <small>[[Francisco De Figueiredo|De Figueiredo]], 2009</small><ref>{{Cite journal |last=De Figueiredo |first=Francisco J. |date=2009-07-21 |title=A new marine clupeoid fish from the Lower Cretaceous of the Sergipe-Alagoas Basin, northeastern Brazil |url=https://biotaxa.org/Zootaxa/article/view/zootaxa.2164.1.2 |journal=Zootaxa |volume=2164 |issue=1 |pages=21–32 |doi=10.11646/zootaxa.2164.1.2 |issn=1175-5334|url-access=subscription }}</ref>
*** Genus †''[[Pseudoellimma|Pseudoellima]]'' <small>De Figueiredo, 2009</small><ref name="De Figueiredo" />
*** Genus †''[[Pseudoellimma|Pseudoellima]]'' <small>De Figueiredo, 2009</small><ref name="De Figueiredo" />
*** Family †[[Cynoclupeidae]] <small>[[Maria Claudia Malabarba|Malabarba]] & [[Fabio Di Dario|Di Dario]], 2017</small><ref name=":0" />
*** Family †[[Cynoclupeidae]] <small>[[Maria Claudia Malabarba|Malabarba]] & [[Fabio Di Dario|Di Dario]], 2017</small><ref name=":0" />
*** Family †[[Garganoclupeidae]] <small>Taverne, 2014</small><ref>{{Cite journal |last=Laan |first=Richard van der |date=2018-10-11 |title=Family-group names of fossil fishes |url=https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/597 |journal=European Journal of Taxonomy |language=en |issue=466 |doi=10.5852/ejt.2018.466 |issn=2118-9773}}</ref>
*** Family [[Spratelloididae]] <small>[[David Starr Jordan|D. S. Jordan]] 1925</small> (dwarf herrings or small round herrings)
*** Family [[Spratelloididae]] <small>[[David Starr Jordan|D. S. Jordan]] 1925</small> (dwarf herrings or small round herrings)
*** Family [[Engraulidae]] <small>[[Theodore Gill|Gill]], 1861</small> (anchovies)
*** Family [[Engraulidae]] <small>[[Theodore Gill|Gill]], 1861</small> (anchovies)
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==References==
==References==
{{Reflist}}
{{Reflist}}
* {{cite journal|last=Sepkoski |first=Jack |title=A compendium of fossil marine animal genera |journal=Bulletins of American Paleontology |volume=363 |pages=1–560 |year=2002 |url=http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class |access-date=2011-05-17 |url-status=dead |archive-url=https://web.archive.org/web/20090220223520/http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class |archive-date=2009-02-20 }}
* {{cite journal|last=Sepkoski |first=Jack |title=A compendium of fossil marine animal genera |journal=Bulletins of American Paleontology |volume=363 |pages=1–560 |year=2002 |url=http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class |access-date=2011-05-17 |archive-url=https://web.archive.org/web/20090220223520/http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class |archive-date=2009-02-20 }}


{{Actinopterygii}}
{{Actinopterygii}}

Latest revision as of 22:37, 27 September 2025

Template:Short description Template:Automatic taxobox

Clupeiformes Template:IPAc-en is the order of ray-finned fish that includes the herring family, Clupeidae, and the anchovy family, Engraulidae and sardines. The group includes many of the most important forage and food fish.

Clupeiformes are physostomes, which means that their gas bladder has a pneumatic duct connecting it to the gut. They typically lack a lateral line, but still have the eyes, fins and scales that are common to most fish, though not all fish have these attributes. They are generally silvery fish with streamlined, spindle-shaped bodies, and they often school. Most species eat plankton which they filter from the water with their gill rakers.[1]

Script error: No such module "anchor". The former order of Isospondyli was subsumed mostly by Clupeiformes,[2] but some isospondylous fishes (isospondyls) were assigned to Osteoglossiformes, Salmoniformes, Cetomimiformes, etc.[3]

Their sister group were the extinct Ellimmichthyiformes, which were dominant throughout much of the Cretaceous and into the Paleogene,[4] and often coexisted with clupeiforms at many known localities. Both groups closely resembled each other morphologically, although the ellimmichthyiformes evolved some highly divergent body plans later in the Cretaceous.

Several fossil clupeiforms are known from the Early Cretaceous of South America that appear to be more closely allied with Clupeioidei over the Denticipitidae. This suggests a very deep divergence within the crown group Clupeiformes that must have occurred during the Early Cretaceous or before.[5][6]

Families

Template:Cladogram The order includes about 405 species in ten families:[7][8][9]

Timeline of genera

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Period = from:-199.6 till:15 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:10 start:-199.6 ScaleMinor = unit:year increment:1 start:-199.6 TimeAxis = orientation:hor AlignBars = justify

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bar:periodtop          
from: -199.6    till: -175.6    color:earlyjurassic    text:Early
from: -175.6    till: -161.2    color:middlejurassic    text:Middle
from: -161.2    till: -145.5    color:latejurassic    text:Late
from: -145.5    till: -99.6    color:earlycretaceous    text:Early
from: -99.6    till: -65.5    color:latecretaceous    text:Late
from: -65.5   till:  -55.8    color:paleocene  text:Paleo.
from: -55.8   till:  -33.9    color:eocene  text:Eo. 
from: -33.9   till:  -23.03    color:oligocene  text:Oligo.            
from: -23.03    till: -5.332    color:miocene    text:Mio.
from: -5.332    till: -2.588    color:pliocene    text:Pl.
from: -2.588    till: -0.0117   color:pleistocene    text:Pl.
from: -0.0117    till: 0    color:holocene    text:H.
bar:eratop       
from: -199.6    till: -145.5    color:jurassic    text:Jurassic
from: -145.5    till: -65.5   color:cretaceous    text:Cretaceous
from: -65.5   till:  -23.03    color:paleogene  text:Paleogene         
from: -23.03    till: -2.588    color:neogene    text:Neogene
from: -2.588    till: 0   color:quaternary    text:Q.

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align:left fontsize:M mark:(line,white) width:5 anchor:till align:left
color:latejurassic bar:NAM1  from:-161.2    till:-149.03 text:Pachythrissops
color:latejurassic bar:NAM2  from:-150.8    till:-149.03 text:Daitingichthys
color:latecretaceous bar:NAM3  from:-99.6    till:-93.5 text:Ornategulum
color:latecretaceous bar:NAM4 from:-99.6    till:-93.5 text:Scombroclupea
color:latecretaceous bar:NAM5  from:-94.3    till:-85.8 text:Leufuichthys
color:latecretaceous bar:NAM6 from:-85.8    till:-83.5 text:Histiothrissa
color:latecretaceous bar:NAM7 from:-70.6    till:-65.5 text:Gasteroclupea
color:eocene bar:NAM6 from:-55.8   till:-48.6 text:Knightia
color:eocene bar:NAM8  from:-55.8    till:-33.9 text:Hacquetia
color:eocene bar:NAM9  from:-55.8    till:0 text:Clupea
color:eocene bar:NAM10  from:-55.8    till:0 text:Harengula
color:eocene bar:NAM11 from:-55.8    till:0 text:Chirocentrus
color:eocene bar:NAM12 from:-48.6    till:0 text:Etrumeus
color:eocene bar:NAM13 from:-48.6    till:0 text:Sardinella
color:eocene bar:NAM14  from:-37.2    till:0 text:Alosa
color:oligocene bar:NAM15 from:-33.9    till:0 text:Opisthonema
color:oligocene bar:NAM16  from:-33.9    till:0 text:Pomolobus
color:oligocene bar:NAM17  from:-28.4    till:0 text:Sardina
color:miocene bar:NAM18  from:-23.03    till:0 text:Stolephorus
color:miocene bar:NAM19 from:-23.03    till:0 text:Anchoa
color:miocene bar:NAM20 from:-23.03    till:0 text:Ilisha
color:miocene bar:NAM21 from:-23.03    till:0 text:Sardinops
color:miocene bar:NAM22 from:-15.97    till:0 text:Pseudohilsa
color:miocene bar:NAM23 from:-11.608    till:-5.332 text:Alisea
color:miocene bar:NAM24 from:-11.608    till:-5.332 text:Etringus
color:miocene bar:NAM25 from:-11.608    till:-5.332 text:Ganoessus
color:miocene bar:NAM26 from:-11.608    till:-5.332 text:Ganolytes
color:miocene bar:NAM27 from:-11.608    till:-5.332 text:Quisque
color:miocene bar:NAM28 from:-11.608    till:-5.332 text:Xyne
color:miocene bar:NAM29 from:-11.608    till:0 text:Engraulis
color:miocene bar:NAM30 from:-11.608    till:0 text:Sarmatella
color:pliocene bar:NAM31 from:-5.332    till:-2.588 text:Austroclupea

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from: -199.6    till: -175.6    color:earlyjurassic    text:Early
from: -175.6    till: -161.2    color:middlejurassic    text:Middle
from: -161.2    till: -145.5    color:latejurassic    text:Late
from: -145.5    till: -99.6    color:earlycretaceous    text:Early
from: -99.6    till: -65.5    color:latecretaceous    text:Late
from: -65.5   till:  -55.8    color:paleocene  text:Paleo.
from: -55.8   till:  -33.9    color:eocene  text:Eo. 
from: -33.9   till:  -23.03    color:oligocene  text:Oligo.            
from: -23.03    till: -5.332    color:miocene    text:Mio.
from: -5.332    till: -2.588    color:pliocene    text:Pl.
from: -2.588    till: -0.0117   color:pleistocene    text:Pl.
from: -0.0117    till: 0    color:holocene    text:H.
bar:era   
from: -199.6    till: -145.5    color:jurassic    text:Jurassic
from: -145.5    till: -65.5   color:cretaceous    text:Cretaceous
from: -65.5   till:  -23.03    color:paleogene  text:Paleogene         
from: -23.03    till: -2.588    color:neogene    text:Neogene
from: -2.588    till: 0   color:quaternary    text:Q.

</timeline>

References

Template:Reflist

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Template:Actinopterygii Template:Diversity of fish Template:Taxonbar Template:Authority control

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