Percidae: Difference between revisions

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The '''Percidae''' are a [[Family (biology)|family]] of [[ray-finned fish]], part of the [[Order (biology)|order]] [[Perciformes]], which are found in fresh and [[brackish]] waters of the [[Northern Hemisphere]]. The majority are [[Nearctic]], but there are also [[Palearctic]] species. The family contains nearly 250 species in 12 genera.<ref name=":0">{{Cite web |title=CAS - Eschmeyer's Catalog of Fishes - Genera/Species by Family/Subfamily |url=https://researcharchive.calacademy.org/research/ichthyology/catalog/SpeciesByFamily.asp |access-date=2025-04-22 |website=researcharchive.calacademy.org}}</ref> The perches and their relatives are in this family; well-known species include the [[walleye]], [[sauger]], [[ruffe]], and three species of [[perch]]. However, small fish known as [[Darter (fish)|darters]] are also a part of this family.  
The '''Percidae''' are a [[Family (biology)|family]] of [[ray-finned fish]], part of the [[Order (biology)|order]] [[Perciformes]], which are found in fresh and [[brackish]] waters of the [[Northern Hemisphere]]. The majority are [[Nearctic]], but there are also [[Palearctic]] species. The family contains nearly 250 species in 12 genera.<ref name=":0">{{Cite web |title=CAS - Eschmeyer's Catalog of Fishes - Genera/Species by Family/Subfamily |url=https://researcharchive.calacademy.org/research/ichthyology/catalog/SpeciesByFamily.asp |access-date=2025-04-22 |website=researcharchive.calacademy.org}}</ref> The perches and their relatives are in this family; well-known species include the [[walleye]], [[sauger]], [[ruffe]], and three species of [[perch]]. However, small fish known as [[Etheostomatinae|darters]] are also a part of this family.  


Despite the widespread and speciose nature of this family, the vast majority of its species diversity is within the darters, which are restricted to eastern North America, and the majority of which have small ranges. Excluding the darters, this family contains only 20 species.<ref name=":0" /> In Europe, percids were likely a more ecologically dominant group in the past than now, with the immigration of [[Cyprinidae|cyprinids]] to Europe during the [[Miocene]] causing a decline in their importance.<ref name=":1">{{Cite journal |last=Kovalchuk |first=Oleksandr M. |last2=and Murray |first2=Alison M. |date=2016-05-03 |title=Late Miocene and Pliocene pikeperches (Teleostei, Percidae) of southeastern Europe |url=https://www.tandfonline.com/doi/full/10.1080/02724634.2016.1100999 |journal=Journal of Vertebrate Paleontology |volume=36 |issue=3 |pages=e1100999 |doi=10.1080/02724634.2016.1100999 |issn=0272-4634|url-access=subscription }}</ref>
Despite the widespread and speciose nature of this family, the vast majority of its species diversity is within the darters, which are restricted to eastern North America, and the majority of which have small ranges. Excluding the darters, this family contains only 20 species.<ref name=":0" /> In Europe, percids were likely a more ecologically dominant group in the past than now, with the immigration of [[Cyprinidae|cyprinids]] to Europe during the [[Miocene]] causing a decline in their importance.<ref name=":1">{{Cite journal |last1=Kovalchuk |first1=Oleksandr M. |last2=and Murray |first2=Alison M. |date=2016-05-03 |title=Late Miocene and Pliocene pikeperches (Teleostei, Percidae) of southeastern Europe |url=https://www.tandfonline.com/doi/full/10.1080/02724634.2016.1100999 |journal=Journal of Vertebrate Paleontology |volume=36 |issue=3 |article-number=e1100999 |doi=10.1080/02724634.2016.1100999 |bibcode=2016JVPal..36E0999K |issn=0272-4634|url-access=subscription }}</ref>


==Characteristics==
==Characteristics==
The family is characterised by having the [[dorsal fin]] split into two<ref name = IFNY>{{cite web | author1 = Kraft C.E | author2 = D.M. Carlson | author3 = M. Carlson | year = 2006  |title = Inland fishes of New York (online) Version4.0 | url = http://www2.dnr.cornell.edu/cek7/nyfish/index.html | publisher = Department of Natural Resources, Cornell University and the New York State Department of Environmental Conservation}}</ref> which are normally separated or have a narrow connection, although this is wider in the genus ''Zingel'',<ref name="Nelson5">{{cite book |author1=J. S. Nelson |url=https://sites.google.com/site/fotw5th/ |title=Fishes of the World |author2=T. C. Grande |author3=M. V. H. Wilson |publisher=Wiley |year=2016 |ISBN=978-1-118-34233-6 |edition=5th |pages=448–450}}</ref> the front section contains the spines and the rear section contains the soft rays.<ref name = IFNY/> The [[anal fin]] contains 1 or 2 spines, if there is a second spine it is typically weak. The [[pelvic fins]] are placed on the [[thorax]] and have a single spine and 5 soft rays. They also have skeletal [[synapomorphy|synapomorphies]].<ref name = Nelson5/> The maximum size attained is {{convert|100|cm|in}} in the [[zander]] (''Sander lucioperca'')<ref name="Fishbase">{{FishBase family|family=Percidae|month=December|year=2019}}</ref> but most of the species in the family are much smaller.<ref name = Nelson5/> Their scales are [[ctenoid]] and their bodies are normally somewhat elongate.<ref name = Britannia>{{cite web | url = https://www.britannica.com/animal/perciform/Form-and-function#ref1044889 | title = Percidae: Form and Function | access-date = 12 September 2020 | publisher = Encyclopaedia Britannica}}</ref>
The family is characterised by having the [[dorsal fin]] split into two<ref name = IFNY>{{cite web | author1 = Kraft C.E | author2 = D.M. Carlson | author3 = M. Carlson | year = 2006  |title = Inland fishes of New York (online) Version4.0 | url = https://www2.dnr.cornell.edu/cek7/nyfish/index.html | publisher = Department of Natural Resources, Cornell University and the New York State Department of Environmental Conservation}}</ref> which are normally separated or have a narrow connection, although this is wider in the genus ''Zingel'',<ref name="Nelson5">{{cite book |author1=J. S. Nelson |url=https://sites.google.com/site/fotw5th/ |title=Fishes of the World |author2=T. C. Grande |author3=M. V. H. Wilson |publisher=Wiley |year=2016 |isbn=978-1-118-34233-6 |edition=5th |pages=448–450}}</ref> the front section contains the spines and the rear section contains the soft rays.<ref name = IFNY/> The [[anal fin]] contains 1 or 2 spines, if there is a second spine it is typically weak. The [[pelvic fins]] are placed on the [[thorax]] and have a single spine and 5 soft rays. They also have skeletal [[synapomorphy|synapomorphies]].<ref name = Nelson5/> The maximum size attained is {{convert|100|cm|in}} in the [[zander]] (''Sander lucioperca'')<ref name="Fishbase">{{FishBase family|family=Percidae|month=December|year=2019}}</ref> but most of the species in the family are much smaller.<ref name = Nelson5/> Their scales are [[ctenoid]] and their bodies are normally somewhat elongate.<ref name = Britannia>{{cite web | url = https://www.britannica.com/animal/perciform/Form-and-function#ref1044889 | title = Percidae: Form and Function | access-date = 12 September 2020 | publisher = Encyclopaedia Britannica}}</ref>


==Systematics==
==Systematics==

Latest revision as of 18:28, 3 October 2025

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The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains nearly 250 species in 12 genera.[1] The perches and their relatives are in this family; well-known species include the walleye, sauger, ruffe, and three species of perch. However, small fish known as darters are also a part of this family.

Despite the widespread and speciose nature of this family, the vast majority of its species diversity is within the darters, which are restricted to eastern North America, and the majority of which have small ranges. Excluding the darters, this family contains only 20 species.[1] In Europe, percids were likely a more ecologically dominant group in the past than now, with the immigration of cyprinids to Europe during the Miocene causing a decline in their importance.[2]

Characteristics

The family is characterised by having the dorsal fin split into two[3] which are normally separated or have a narrow connection, although this is wider in the genus Zingel,[4] the front section contains the spines and the rear section contains the soft rays.[3] The anal fin contains 1 or 2 spines, if there is a second spine it is typically weak. The pelvic fins are placed on the thorax and have a single spine and 5 soft rays. They also have skeletal synapomorphies.[4] The maximum size attained is Template:Convert in the zander (Sander lucioperca)[5] but most of the species in the family are much smaller.[4] Their scales are ctenoid and their bodies are normally somewhat elongate.[6]

Systematics

The following classification is based on Eschmeyer's Catalog of Fishes (2025):[7]

Under a former treatment by the 5th Edition of Fishes of the World, Gymnocephalus was placed in its own subfamily, Acerinae, as was Percarina in Percarininae.[4][5]

Although the family likely underwent an evolutionary radiation during the Paleogene, the earliest definitive fossil remains of percids are of modern Sander from the Middle Miocene of Canada. †Vixperca Peña Zarzuelo, 1991 from the Middle Eocene of Spain has been suggested to be the earliest known percid, but its remains are fragmentary and show closer similarities to lates perches than to true perches.[2]

References

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