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	<title>Isothrix - Revision history</title>
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	<updated>2026-05-12T15:52:34Z</updated>
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		<title>imported&gt;Dimadick: /* References */</title>
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		<updated>2025-05-28T07:52:09Z</updated>

		<summary type="html">&lt;p&gt;&lt;span class=&quot;autocomment&quot;&gt;References&lt;/span&gt;&lt;/p&gt;
&lt;p&gt;&lt;b&gt;New page&lt;/b&gt;&lt;/p&gt;&lt;div&gt;{{Short description|Genus of mammals belonging to the spiny rat family of rodents}}&lt;br /&gt;
{{automatic taxobox&lt;br /&gt;
| fossil_range = [[Pleistocene]] - Recent&lt;br /&gt;
| image = Isothrix bistriata 74499273 (cropped).jpg&lt;br /&gt;
| image_caption = &amp;#039;&amp;#039;[[Isothrix bistriata]]&amp;#039;&amp;#039;&lt;br /&gt;
| display_parents = 2&lt;br /&gt;
| taxon = Isothrix&lt;br /&gt;
| authority = [[Johann Andreas Wagner|Wagner]], 1845&lt;br /&gt;
| type_species = &amp;#039;&amp;#039;[[Isothrix bistriata]]&amp;#039;&amp;#039;&lt;br /&gt;
| type_species_authority = [[Johann Andreas Wagner|Wagner]], 1845&lt;br /&gt;
| subdivision_ranks = Species&lt;br /&gt;
| subdivision =&lt;br /&gt;
&amp;#039;&amp;#039;[[Isothrix barbarabrownae|I. barbarabrownae]]&amp;#039;&amp;#039;&amp;lt;br&amp;gt;&lt;br /&gt;
&amp;#039;&amp;#039;[[Isothrix bistriata|I. bistriata]]&amp;#039;&amp;#039;&amp;lt;br&amp;gt;&lt;br /&gt;
&amp;#039;&amp;#039;[[Isothrix negrensis|I. negrensis]]&amp;#039;&amp;#039;&amp;lt;br&amp;gt;&lt;br /&gt;
&amp;#039;&amp;#039;[[Isothrix pagurus|I. pagurus]]&amp;#039;&amp;#039;&amp;lt;br&amp;gt;&lt;br /&gt;
&amp;#039;&amp;#039;[[Isothrix sinnamariensis|I. sinnamariensis]]&amp;#039;&amp;#039;&lt;br /&gt;
| synonyms =&lt;br /&gt;
&amp;#039;&amp;#039;Lasiuromys&amp;#039;&amp;#039; &amp;lt;small&amp;gt;Deville, 1852&amp;lt;/small&amp;gt;&lt;br /&gt;
}}&lt;br /&gt;
&lt;br /&gt;
The &amp;#039;&amp;#039;&amp;#039;toros&amp;#039;&amp;#039;&amp;#039; or &amp;#039;&amp;#039;&amp;#039;brush-tailed rats&amp;#039;&amp;#039;&amp;#039;, genus &amp;#039;&amp;#039;&amp;#039;&amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039;&amp;#039;&amp;#039;&amp;#039;, are a group of [[Echimyidae|spiny rat]]s&amp;lt;ref&amp;gt;{{MSW3|id=13400433}}&amp;lt;/ref&amp;gt; found in [[tropical]] [[South America]], particularly in the [[Amazon Basin]].&amp;lt;ref name=&amp;quot;Nowak1999&amp;quot;&amp;gt;Nowak, Ronald M. 1999. &amp;#039;&amp;#039;Walker&amp;#039;s Mammals of the World&amp;#039;&amp;#039;, 6th edition. Johns Hopkins University Press, 1936 pp.&amp;amp;nbsp;{{ISBN|0-8018-5789-9}}&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
==Description==&lt;br /&gt;
Toros look like large [[rat]]s with soft fur on the body and long [[guard hair]]s on the scaly tail.&amp;lt;ref name=&amp;quot;Emmons2005&amp;quot;&amp;gt;Emmons, L.H. 2005. A Revision of the Genera of Arboreal Echimyidae (Rodentia: Echimyidae, Echimyinae), With Descriptions of Two New Genera. pp.&amp;amp;nbsp;247–310 in Lacey, E.A. &amp;amp; Myers, P. 2005. Mammalian Diversification: From Chromosomes to Phylogeography (A Celebration of the Career of James L. Patton). University of California Publications in Zoology.&amp;lt;/ref&amp;gt;  Head and body is 18-27.5&amp;amp;nbsp;cm and tail is 17–30&amp;amp;nbsp;cm.&amp;lt;ref name=&amp;quot;Nowak1999&amp;quot;/&amp;gt; Weight is 320-570&amp;amp;nbsp;grams.&lt;br /&gt;
&lt;br /&gt;
==Natural history==&lt;br /&gt;
These animals appear to be [[arboreal]], based on the shape of their hind feet.&amp;lt;ref name=&amp;quot;Nowak1999&amp;quot;/&amp;gt; They are thought to spend the day in holes in the ground near trees.&lt;br /&gt;
&lt;br /&gt;
==Systematics==&lt;br /&gt;
The etymology of the genus name &amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039; corresponds to the two [[ancient greek]] words {{wikt-lang|grc|ἴσος}} ({{grc-transl|ἴσος}}), meaning &amp;quot;of equal length&amp;quot;, and {{wikt-lang|grc|θρίξ}}, {{wikt-lang|grc|τρίχα|τριχός}} ({{grc-transl|θρίξ, τριχός}}), meaning &amp;quot;hair&amp;quot;.&amp;lt;ref name=&amp;quot;BaillyBook&amp;quot;&amp;gt;{{Cite book |title=Abrégé du dictionnaire grec français |last=Bailly |first=Anatole |date=1981-01-01 |publisher=Hachette |isbn=978-2010035289 |location=Paris |oclc=461974285 }}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;BaillyWeb&amp;quot;&amp;gt;{{Cite web |url=http://www.tabularium.be/bailly/ |title=Greek-french dictionary online |last=Bailly |first=Anatole |website=www.tabularium.be |access-date=December 23, 2017}}&amp;lt;/ref&amp;gt; The etymology of the genus name [[Synonym (taxonomy)|synonym]] &amp;#039;&amp;#039;Lasiuromys&amp;#039;&amp;#039; corresponds to the three [[ancient greek]] words {{wikt-lang|grc|λάσιος}} ({{grc-transl|λάσιος}}), meaning &amp;quot;furry, with tufted hair&amp;quot;, {{wikt-lang|grc|οὐρά}} ({{grc-transl|οὐρά}}), meaning &amp;quot;animal tail&amp;quot;, and {{wikt-lang|grc|μῦς}} ({{grc-transl|μῦς}}), meaning &amp;quot;mouse, rat&amp;quot;.&amp;lt;ref name=&amp;quot;BaillyBook&amp;quot;/&amp;gt;&amp;lt;ref name=&amp;quot;BaillyWeb&amp;quot;/&amp;gt;&lt;br /&gt;
&lt;br /&gt;
There are currently five [[species]] recognized in the [[genus]] &amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039;:&amp;lt;ref name=&amp;quot;MSW3_Isothrix&amp;quot;&amp;gt;{{MSW3 Hystricognathi | id = 13400433 | page = | heading = Genus &amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039; }}&amp;lt;/ref&amp;gt;&lt;br /&gt;
* &amp;#039;&amp;#039;[[Isothrix barbarabrownae]]&amp;#039;&amp;#039;&amp;lt;ref name = &amp;quot;New_Scientist&amp;quot;&amp;gt;{{cite web| title = New squirrel-like rodent discovered in Peru | publisher = [[New Scientist]] | date = 2007-01-24 | url = https://www.newscientist.com/article.ns?id=dn11022 | access-date = 2011-11-23}}&amp;lt;/ref&amp;gt;- Barbara Brown&amp;#039;s Brush-tailed Rat&lt;br /&gt;
* &amp;#039;&amp;#039;[[Isothrix bistriata]]&amp;#039;&amp;#039; - Yellow-crowned brush-tailed rat&lt;br /&gt;
* &amp;#039;&amp;#039;[[Isothrix negrensis]]&amp;#039;&amp;#039; - Rio Negro brush-tailed rat&lt;br /&gt;
* &amp;#039;&amp;#039;[[Isothrix pagurus]]&amp;#039;&amp;#039; - Plain brush-tailed rat&lt;br /&gt;
* &amp;#039;&amp;#039;[[Isothrix sinnamariensis]]&amp;#039;&amp;#039; - Sinnamary brush-tailed rat&lt;br /&gt;
&lt;br /&gt;
There does appear to be good support for the [[monophyly]] of three species found in this genus.&amp;lt;ref name=&amp;quot;Emmons2005&amp;quot;/&amp;gt;&lt;br /&gt;
&lt;br /&gt;
==Phylogeny==&lt;br /&gt;
[[File:Echimyidae_phylogeny.jpg|thumb]]&lt;br /&gt;
&amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039; is a member of the [[Echimyini]] clade of [[arboreal]] Echimyidae rodents.&amp;lt;ref name=&amp;quot;Fabre2016&amp;quot;&amp;gt;{{Cite book|title=Handbook of the Mammals of the World. Vol 6. Lagomorphs and Rodents I|last1=Fabre|first1=Pierre-Henri|last2=Patton|first2=James L.|author2-link=James L. Patton|last3=Leite|first3=Yuri L. R.|year=2016|isbn=978-84-941892-3-4|editor-last1=Wilson|editor-first1=Don E.|editor-last2=Lacher|editor-first2=Thomas E. Jr|editor-last3=Mittermeier|editor-first3=Russell A.|publisher=Lynx Edicions|location=Barcelona|pages=552–641|chapter=Family Echimyidae (hutias, South American spiny-rats and coypu)|title-link=Handbook of the Mammals of the World}}&amp;lt;/ref&amp;gt; Although tentatively considered an [[Echimyinae|echimyine]], it has been suggested that &amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039; may not be particularly closely related to other members of its subfamily.&amp;lt;ref name=&amp;quot;Emmons2005&amp;quot;/&amp;gt; This is confirmed by molecular phylogenies in which &amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039; appears as a distant relative of the three clades of [[Echimyini]]: (i) &amp;#039;&amp;#039;Echimys&amp;#039;&amp;#039;, &amp;#039;&amp;#039;Phyllomys&amp;#039;&amp;#039;, &amp;#039;&amp;#039;Makalata&amp;#039;&amp;#039;, &amp;#039;&amp;#039;Pattonomys&amp;#039;&amp;#039;, and &amp;#039;&amp;#039;Toromys&amp;#039;&amp;#039; ; (ii) the bamboo rats &amp;#039;&amp;#039;Dactylomys&amp;#039;&amp;#039;, &amp;#039;&amp;#039;Olallamys&amp;#039;&amp;#039;, &amp;#039;&amp;#039;Kannabateomys&amp;#039;&amp;#039; together with &amp;#039;&amp;#039;Diplomys&amp;#039;&amp;#039; and &amp;#039;&amp;#039;Santamartamys&amp;#039;&amp;#039; ; and (iii) the arboreal eumysopines &amp;#039;&amp;#039;Mesomys&amp;#039;&amp;#039; and &amp;#039;&amp;#039;Lonchothrix&amp;#039;&amp;#039;.&amp;lt;ref name=&amp;quot;Fabre2017&amp;quot;&amp;gt;{{Cite journal|last1=Fabre|first1=Pierre-Henri|last2=Upham|first2=Nathan S.|last3=Emmons|first3=Louise H.|author-link3=Louise H. Emmons|last4=Justy|first4=Fabienne|last5=Leite|first5=Yuri L. R.|last6=Loss|first6=Ana Carolina|last7=Orlando|first7=Ludovic|last8=Tilak|first8=Marie-Ka|last9=Patterson|first9=Bruce D.|last10=Douzery|first10=Emmanuel J. P.|date=2017-03-01|title=Mitogenomic Phylogeny, Diversification, and Biogeography of South American Spiny Rats|journal=Molecular Biology and Evolution|volume=34|issue=3|pages=613–633|doi=10.1093/molbev/msw261|pmid=28025278|issn=0737-4038|doi-access=free|url=https://hal.umontpellier.fr/hal-03079822/file/msw261.pdf}}&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
The taxon &amp;#039;&amp;#039;[[Callistomys pictus]]&amp;#039;&amp;#039; was previously member of the genus &amp;#039;&amp;#039;Isothrix&amp;#039;&amp;#039;, but most authors considered it distinct enough to warrant a separate genus.&amp;lt;ref name=&amp;quot;MSW3_Callistomys&amp;quot;&amp;gt;{{MSW3 Hystricognathi | id = 13400423 | page = | heading = Genus &amp;#039;&amp;#039;Callistomys&amp;#039;&amp;#039;}}&amp;lt;/ref&amp;gt; This distinctness has been confirmed by DNA comparative studies in which &amp;#039;&amp;#039;Callistomys&amp;#039;&amp;#039; even appears to branch with members of a distinct tribe (i.e., [[Myocastorini]]) of the subfamily Echimyinae.&amp;lt;ref name=&amp;quot;Loss2014&amp;quot;/&amp;gt;&amp;lt;ref name=&amp;quot;Fabre2016&amp;quot;/&amp;gt;&amp;lt;ref name=&amp;quot;Fabre2017&amp;quot;/&amp;gt; The arboreal genera of the subfamily Echimyinae therefore constitute a [[polyphyletic]] assemblage.&lt;br /&gt;
&lt;br /&gt;
{{cladogram&lt;br /&gt;
   |title=Genus-level cladogram of the Echimyini.&lt;br /&gt;
   |caption=The cladogram has been reconstructed from mitochondrial and nuclear DNA characters.&amp;lt;ref name=&amp;quot;Lara1996&amp;quot;&amp;gt;{{Cite journal|last1=Lara|first1=Márcia C.|last2=Patton|first2=James L.|author-link2=James L. Patton|last3=da Silva|first3=Maria Nazareth F.|year=1996|title=The Simultaneous Diversification of South American Echimyid Rodents (Hystricognathi) Based on Complete Cytochrome &amp;#039;&amp;#039;b&amp;#039;&amp;#039; Sequences|journal=Molecular Phylogenetics and Evolution|volume=5|issue=2|pages=403–413|doi=10.1006/mpev.1996.0035|pmid=8728398|bibcode=1996MolPE...5..403L }}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Leite2002&amp;quot;&amp;gt;{{Cite journal|last1=Leite|first1=Yuri L. R.|last2=Patton|first2=James L.|author-link2=James L. Patton|year=2002|title=Evolution of South American spiny rats (Rodentia, Echimyidae): the star-phylogeny hypothesis revisited|journal=Molecular Phylogenetics and Evolution|volume=25|issue=3|pages=455–464|doi=10.1016/s1055-7903(02)00279-8|pmid=12450750 |bibcode=2002MolPE..25..455L }}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Galewski2005&amp;quot;&amp;gt;{{Cite journal|last1=Galewski|first1=Thomas|last2=Mauffrey|first2=Jean-François|last3=Leite|first3=Yuri L. R.|last4=Patton|first4=James L.|last5=Douzery|first5=Emmanuel J. P.|year=2005|title=Ecomorphological diversification among South American spiny rats (Rodentia; Echimyidae): a phylogenetic and chronological approach|journal=Molecular Phylogenetics and Evolution|volume=34|issue=3|pages=601–615|doi=10.1016/j.ympev.2004.11.015|pmid=15683932|bibcode=2005MolPE..34..601G }}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Upham2012&amp;quot;&amp;gt;{{Cite journal|last1=Upham|first1=Nathan S.|last2=Patterson|first2=Bruce D.|year=2012|title=Diversification and biogeography of the Neotropical caviomorph lineage Octodontoidea (Rodentia: Hystricognathi)|journal=Molecular Phylogenetics and Evolution|volume=63|issue=2|pages=417–429|doi=10.1016/j.ympev.2012.01.020|pmid=22327013|bibcode=2012MolPE..63..417U }}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Fabre2013&amp;quot;&amp;gt;{{Cite journal|last1=Fabre|first1=Pierre-Henri|last2=Galewski|first2=Thomas|last3=Tilak|first3=Marie-ka|last4=Douzery|first4=Emmanuel J. P.|date=2013-03-01|title=Diversification of South American spiny rats (Echimyidae): a multigene phylogenetic approach|journal=Zoologica Scripta|language=en|volume=42|issue=2|pages=117–134|doi=10.1111/j.1463-6409.2012.00572.x|issn=1463-6409}}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Loss2014&amp;quot;&amp;gt;{{Cite journal|last1=Loss|first1=Ana|last2=Moura|first2=Raquel T.|last3=Leite|first3=Yuri L. R.|date=2014|title=Unexpected phylogenetic relationships of the painted tree rat &amp;#039;&amp;#039;Callistomys pictus&amp;#039;&amp;#039; (Rodentia: Echimyidae)|url=http://www.naturezaonline.com.br/natureza/conteudo/pdf/05_LossACetal_132-136.pdf|journal=Natureza on Line|volume=12|pages=132–136|access-date=2017-12-22|archive-date=2021-06-23|archive-url=https://web.archive.org/web/20210623231029/http://www.naturezaonline.com.br/natureza/conteudo/pdf/05_LossACetal_132-136.pdf|url-status=dead}}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Fabre2014&amp;quot;&amp;gt;{{Cite journal|last1=Fabre|first1=Pierre-Henri|last2=Vilstrup|first2=Julia T.|last3=Raghavan|first3=Maanasa|last4=Der Sarkissian|first4=Clio|last5=Willerslev|first5=Eske|last6=Douzery|first6=Emmanuel J. P.|last7=Orlando|first7=Ludovic|date=2014-07-01|title=Rodents of the Caribbean: origin and diversification of hutias unravelled by next-generation museomics|journal=Biology Letters|language=en|volume=10|issue=7|pages=20140266|doi=10.1098/rsbl.2014.0266|pmid=25115033|issn=1744-9561|pmc=4126619}}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Upham2015&amp;quot;&amp;gt;{{Cite book|title=Biology of caviomorph rodents: diversity and evolution|last1=Upham|first1=Nathan S.|last2=Patterson|first2=Bruce D.|publisher=SAREM Series A, Mammalogical Research — Sociedad Argentina para el Estudio de los Mamíferos|year=2015|editor-last1=Vassallo|editor-first1=Aldo Ivan|location=Buenos Aires|pages=63–120|chapter=Evolution of Caviomorph rodents: a complete phylogeny and timetree for living genera|editor-last2=Antenucci|editor-first2=Daniel}}&amp;lt;/ref&amp;gt;&amp;lt;ref name=&amp;quot;Fabre2016&amp;quot;/&amp;gt;&amp;lt;ref name=&amp;quot;Fabre2017&amp;quot;/&amp;gt;&lt;br /&gt;
   |align=center&lt;br /&gt;
   |clades={{Cladogram of Echimyini genera}}&lt;br /&gt;
}}&lt;br /&gt;
&lt;br /&gt;
==References==&lt;br /&gt;
{{Reflist}}&lt;br /&gt;
&lt;br /&gt;
{{Echimyidae nav}}&lt;br /&gt;
{{Taxonbar|from=Q911355}}&lt;br /&gt;
&lt;br /&gt;
[[Category:Isothrix| ]]&lt;br /&gt;
[[Category:Echimyini]]&lt;br /&gt;
[[Category:Rodent genera]]&lt;br /&gt;
[[Category:Taxa named by Johann Andreas Wagner]]&lt;/div&gt;</summary>
		<author><name>imported&gt;Dimadick</name></author>
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