View source for Notosuchia

{{Short description|Extinct suborder of reptiles}}
{{Automatic taxobox
| fossil_range = [[Middle Jurassic]] – [[Messinian|Late Miocene]]/[[Early Pliocene]], {{fossil range|167|5|ref=<ref name=PeerJRazanandrongobe>Dal Sasso C, Pasini G, Fleury G, Maganuco S. (2017) Razanandrongobe sakalavae, a gigantic mesoeucrocodylian from the Middle Jurassic of Madagascar, is the oldest known notosuchian. PeerJ 5:e3481 https://doi.org/10.7717/peerj.3481</ref>}}
| image = August 1, 2012 - Simosuchus on Display at the Royal Ontario Museum.jpg
| image_caption = Mounted skeleton of the notosuchian ''[[Simosuchus clarki]]'' in the [[Royal Ontario Museum]]
| display_parents = 3
| taxon = Notosuchia
| authority = Gasparini, 1971
| subdivision_ranks = Families and genera
| subdivision = *{{extinct}}[[Uruguaysuchidae]]
*{{extinct}}'''Peirosauria''' <small>Leardi ''et al.'', 2024</small><ref>{{Cite journal |last1=Leardi |first1=J. M. |last2=Pol |first2=D. |last3=Montefeltro |first3=F. |last4=Marinho |first4=T. S. |last5=Ruiz |first5=J. V. |last6=Bravo |first6=G. G. |last7=Pinheiro |first7=A. E. P. |last8=Godoy |first8=P. L. |last9=Nicholl |first9=C. S. C. |last10=Lecuona |first10=A. |last11=Larsson |first11=H. C. E. |title=Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes) |year=2024 |journal=Bulletin of Phylogenetic Nomenclature |volume=1 |issue=3 |pages=44–82 |doi=10.11646/bpn.1.3.2 |doi-access=free }}</ref>
**{{extinct}}[[Itasuchidae]]
**{{extinct}}[[Mahajangasuchidae]]
**{{extinct}}[[Peirosauridae]]
*{{extinct}}'''[[Ziphosuchia]]''' <small>Ortega ''et al.'', 2000</small>
**{{extinct}}''[[Libycosuchus]]''
**{{extinct}}''[[Simosuchus]]''
**{{extinct}}[[Candidodontidae]]
**{{extinct}}'''Xenodontosuchia''' <small>Ruiz ''et al.'', 2021</small><ref>{{cite journal| vauthors = Ruiz JV, Bronzati M, Ferreira GS, Martins KC, Queiroz MV, Langer MC, Montefeltro FC |title=A new species of ''Caipirasuchus'' (Notosuchia, Sphagesauridae) from the Late Cretaceous of Brazil and the evolutionary history of Sphagesauria |year=2021 |journal=Journal of Systematic Palaeontology |volume=19 |issue=4 |pages=265–287 |doi=10.1080/14772019.2021.1888815 |bibcode=2021JSPal..19..265R |s2cid=235172623 |url=https://www.researchgate.net/publication/351109083 |access-date=29 April 2022 |via=ResearchGate}}</ref>
***{{extinct}}''[[Chimaerasuchus]]''
***{{extinct}}''[[Comahuesuchus]]''
***{{extinct}}''[[Razanandrongobe]]''?
***{{extinct}}'''[[Sebecosuchia]]'''
***{{extinct}}'''Sphagesauria''' <small>Ruiz ''et al.'', 2021</small><ref>{{cite journal| vauthors = Ruiz JV, Bronzati M, Ferreira GS, Martins KC, Queiroz MV, Langer MC, Montefeltro FC |title=A new species of ''Caipirasuchus'' (Notosuchia, Sphagesauridae) from the Late Cretaceous of Brazil and the evolutionary history of Sphagesauria |year=2021 |journal=Journal of Systematic Palaeontology |volume=19 |issue=4 |pages=265–287 |doi=10.1080/14772019.2021.1888815 |bibcode=2021JSPal..19..265R |s2cid=235172623 |url=https://www.researchgate.net/publication/351109083 |access-date=29 April 2022 |via=ResearchGate}}</ref>
****{{extinct}}''[[Brasileosaurus]]''
****{{extinct}}''[[Coringasuchus]]''
****{{extinct}}''[[Coronelsuchus]]''
****{{extinct}}''[[Eptalofosuchus]]''
****{{extinct}}''[[Llanosuchus]]''
****{{extinct}}''[[Mariliasuchus]]''
****{{extinct}}''[[Notosuchus]]''
****{{extinct}}[[Sphagesauridae]]
}}

'''Notosuchia''' is a [[clade]] of primarily [[Gondwana]]n [[mesoeucrocodylia]]n [[Crocodylomorpha|crocodylomorph]]s that lived during the [[Jurassic]] and [[Cretaceous]]. Some [[phylogeny|phylogenies]] recover [[Sebecosuchia]] as a [[clade]] within Notosuchia, others as a [[sister group]] (see [[#Phylogeny|below]]); if Sebecosuchia is included within Notosuchia its existence is pushed into the [[Late Miocene]] or [[Zanclean|Early Pliocene]], about 7 to 4.5 million years ago.<ref>{{Cite journal |last=Viñola López |first=Lázaro W. |last2=Velez-Juarbe |first2=Jorge |last3=Münch |first3=Philippe |last4=Almonte Milan |first4=Juan N. |last5=Antoine |first5=Pierre-Olivier |last6=Marivaux |first6=Laurent |last7=Jimenez-Vasquez |first7=Osvaldo |last8=Bloch |first8=Jonathan |date=2025 |title=A South American sebecid from the Miocene of Hispaniola documents the presence of apex predators in early West Indies ecosystems |url=https://pmc.ncbi.nlm.nih.gov/articles/PMC12040450/ |journal=Proceedings. Biological Sciences |volume=292 |issue=2045 |pages=20242891 |doi=10.1098/rspb.2024.2891 |issn=1471-2954 |pmc=12040450 |pmid=40300627}}</ref> [[Fossil]]s have been found from [[South America]], [[Africa]], [[Asia]], and [[Europe]]. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory (''[[Chimaerasuchus]]''), omnivory (''[[Simosuchus]]''), and terrestrial hypercarnivory (''[[Baurusuchus]]''). It included many members with highly derived traits unusual for crocodylomorphs, including [[Mammal|mammal-like]] teeth, flexible bands of shield-like body armor similar to those of [[armadillo]]s (''[[Armadillosuchus]]''), and possibly fleshy cheeks and [[Pig|pig-like]] snouts (''[[Notosuchus]]''). The clade was first named in 1971 by [[Zulma Gasparini]] and has since undergone many phylogenetic revisions.<ref name=GZ71>{{cite journal |last=Gasparini |first=Z. |year=1971 |title=Los Notosuchia del Cretácico de América del Sur como un nuevo Infraorden de los Mesosuchia (Crocodilia) |journal=Ameghiniana |volume=8 |pages=83–103}}</ref>

==Description==
Notosuchians were generally small, with slender bodies and erect limbs. The most distinctive characteristics are usually seen in the skull. Notosuchian skulls are generally short and deep. While most are relatively narrow, some are very broad. ''[[Simosuchus]]'' has a broadened skull and jaw that resembles a [[pug]], while ''[[Anatosuchus]]'' has a broad, flat snout like that of a [[duck]].

The teeth vary greatly between different genera. Many have [[heterodont]] dentitions that vary in shape across the jaw. Often, there are large canine-like teeth protruding from the front of the mouth and broader molar-like teeth in the back. Some genera, such as ''[[Yacarerani]]'' and ''[[Pakasuchus]]'', have extremely [[Mammal|mammal-like]] teeth. Their molars are complex and [[multicuspid]], and are able to [[Occlusion (dentistry)|occlude]] or fit with one another. Some forms such as ''[[Malawisuchus]]'' had jaw joints that enabled them to move the jaw back and forth in a shearing motion rather than just up and down.

A derived group of notosuchians, the [[baurusuchid]]s differ considerably from other forms. They are very large in comparison to other notosuchians and are exclusively carnivorous. Baurusuchids have deep skulls and prominent canine-like teeth.

Recent research found ''[[Araripesuchus wegeneri]]'', ''[[Armadillosuchus arrudai]]'', ''[[Baurusuchus]]'', ''[[Iberosuchus macrodon]]'', and ''[[Stratiotosuchus maxhechti]]'' were ectothermic organisms<ref>{{cite journal |last=Cubo |first=J. |author2=Aubier, P. |author3= Faure-Brac, M. G. |author4= Martet, G. |author5= Pellarin, R. |author6= Pelletan, I. |author7= Sena, M. V. A. |year=2022 |title=Paleohistological inferences of thermometabolic regimes inNotosuchia (Pseudosuchia: Crocodylomorpha) revisited |journal=Paleobiology |volume=49 |issue=2 |pages=1–11 |doi=10.1017/pab.2022.28|doi-access=free }}</ref>

==Classification==
===Taxonomy===
====Genera====
The evolutionary interrelationships of Notosuchia are in flux, but the following genera are generally considered notosuchians:

{| class="wikitable sortable"
! Genus
! Age
! Location
! Unit
! class="unsortable" | Notes
! class="unsortable" | Images
|-
| ''[[Adamantinasuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]–[[Santonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
| A [[carnivore]] with a very short, high skull and large eye sockets
| [[File:Adamantinasuchus BW.jpg|125px]]
|-
| ''[[Anatosuchus]]''
| data-sort-value="-121.4 Ma" | [[Aptian]]–[[Albian]]
| {{Flag|Niger}}
| [[Tegama Group]]
| A small notosuchian under {{convert|1|m|ft}} long with a duck-like snout
| [[File:Anatosuchus.jpg|125px]]
|-
| ''[[Araripesuchus]]''
| data-sort-value="-113 Ma" | [[Albian]]–[[Maastrichtian]]
| {{ubl
 | {{flag|Madagascar}}
 | {{flag|Niger}}
 | {{flag|Brazil}}
 | {{flag|Argentina}}
 }}
| {{ubl
 | [[Maevarano Formation]]
 | [[Echkar Formation]]
 | [[Santana Formation]]
 | [[Candeleros Formation]]
 }}
| Six species are known, the most of any notosuchian
| [[File:Araripesuchus wegeneri.jpg|125px]]
|-
| ''[[Armadillosuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]–[[Santonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
| A sphagesaurid with [[armadillo]]-like armor shields.
| [[File:Armadillosuchus.jpg|125px]]
|-
| ''[[Baurusuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
| A large [[hypercarnivore]] {{convert|3.5|to|4|m|ft}} in length
| [[Image:Baurusuchus BW.jpg|125px]]
|-
| ''[[Caipirasuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]–[[Santonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
|
|
|-
| ''[[Campinasuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]–[[Santonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
|
|
|-
| ''[[Candidodon]]''
| data-sort-value="-113 Ma" | [[Albian]]
| {{flag|Brazil}}
| [[Itapecuru Formation]]
|
|
|-
| ''[[Chimaerasuchus]]''
| data-sort-value="-121.4 Ma" | [[Aptian]]–[[Albian]]
| {{flag|China}}
| [[Wulong Formation]]
| The first notosuchian found with [[heterodont]] teeth, thought to be a [[herbivore]]
|
|-
| ''[[Comahuesuchus]]''
| data-sort-value="-86.3 Ma" | [[Santonian]]
| {{flag|Argentina}}
| [[Bajo de la Carpa Formation]]
|
| [[File:Comahuesuchus BW.jpg|125px]]
|-
|
''[[Cynodontosuchus]]''
| data-sort-value="-89.8 Ma" | [[Coniacian]]–[[Santonian]]
| {{flag|Argentina}}
| {{ubl
 | [[Bajo de la Carpa Formation]]
 | [[Pichi Picun Leufu Formation]]
 }}
|
|
|-
| ''[[Libycosuchus]]''
| data-sort-value="-100.5 Ma" | [[Cenomanian]]
| {{ubl
 | {{flag|Egypt}}
 | {{flag|Nigeria}}
 }}
| [[Bahariya Formation]]
|
|
|-
| ''[[Malawisuchus]]''
| data-sort-value="-145 Ma" | [[Early Cretaceous]]
| {{flag|Malawi}}
|
| A possible burrower that could move its jaw back and forth while eating
|
|-
| ''[[Mariliasuchus]]''
| data-sort-value="-83.6 Ma" | [[Campanian]]–[[Maastrichtian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
|
| [[File:Mariliasuchus BW.jpg|125px]]
|-
| ''[[Morrinhosuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]–[[Santonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
|
| [[File:Morrinhosuchus.jpg|125px]]
|-
| ''[[Notosuchus]]''
| data-sort-value="-89.8 Ma" | [[Coniacian]]–[[Santonian]]
| {{flag|Argentina}}
| [[Bajo de la Carpa Formation]]
| A notosuchian that may have had a pig-like snout
| [[File:Notosuchus BW.jpg|125px]]
|-
| ''[[Pakasuchus]]''
| data-sort-value="-113 Ma" | [[Albian]]
| {{flag|Tanzania}}
|
| A notosuchian with very complex, mammal-like heterodont teeth.
| [[File:Pakasuchus.jpg|125px]]
|-
| ''[[Pissarrachampsa]]''
| data-sort-value="-83.6 Ma" | [[Campanian]]–[[Maastrichtian]]
| {{flag|Brazil}}
| [[Vale do Rio do Peixe Formation]]
|
| |[[File:Pissarrachampsa sera.png|125px]]
|-
| ''[[Razanandrongobe]]''
| data-sort-value="-168.2 Ma" | [[Middle Jurassic]] ([[Bathonian]])
| {{flag|Madagascar}}
| [[Sakaraha Formation]]
| The earliest known member of the group.<ref name=PeerJRazanandrongobe />
|
|-
| ''[[Simosuchus]]''
| data-sort-value="-72.1 Ma" | [[Maastrichtian]]
| {{flag|Madagascar}}
|
| A broad-snouted [[omnivore]] with clove-shaped teeth
| [[File:Simosuchus clarki skull.jpg|125px]]
|-
| ''[[Sphagesaurus]]''
| data-sort-value="-100.5 Ma" | [[Late Cretaceous]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
| An omnivorous notosuchian
|
|-
| ''[[Stratiotosuchus]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]–[[Santonian]]
| {{flag|Brazil}}
| [[Adamantina Formation]]
|
| [[File:Stratiotosuchus maxhechti.jpg|125px]]
|-
| ''[[Uruguaysuchus]]''
| data-sort-value="-86.3 Ma" | [[Santonian]]–[[Campanian]]
| {{flag|Uruguay}}
|
|
| [[File:Uruguaysuchus BW.jpg|125px]]
|-
| ''[[Wargosuchus]]''
| data-sort-value="-86.3 Ma" | [[Santonian]]
| {{flag|Argentina}}
| [[Bajo de la Carpa Formation]]
|
|
|-
| ''[[Yacarerani]]''
| data-sort-value="-93.9 Ma" | [[Turonian]]-[[Santonian]]
| {{flag|Bolivia}}
| [[Cajones Formation]]
| A notosuchian with rabbit-like [[incisors]] found in association with a probable nest
| [[File:Yacarerani.jpg|125px]]
|}

===Phylogeny===
{{infobox
|title=[[Cladistics#Cladograms|Cladograms]] of ''Notosuchia''
|bodystyle = background:transparent;
|datastyle = text-align:left;
|data1=
{{Cladogram|title='''Ortega et al., 2000'''<ref name=OBC00/>
|clades={{clade| style=font-size:85%;line-height:80%
|label1='''Notosuchia'''
|1={{clade
   |1=''[[Notosuchus]]''
   |2={{clade
      |1=''[[Libycosuchus]]''
      |2={{clade
         |1=''[[Baurusuchus]]''
         |2={{clade
            |1=''[[Iberosuchus]]''
            |2={{clade
               |1=''[[Sebecus]]''
               |2={{clade
                  |1=Itaborai crocodile
                  |2=''[[Bretesuchus]]''
}} }} }} }} }} }} }}
}}
|data2=
{{Cladogram|title='''Pol, 2003'''<ref name=CB97/>
|clades={{clade| style=font-size:85%;line-height:80%
|label1='''Notosuchia'''
|1={{clade
   |1=''[[Uruguaysuchus]]''
   |2={{clade
      |1=''[[Simosuchus]]''
      |2={{clade
         |1={{clade
            |1=''[[Malawisuchus]]''
            |2=''[[Candidodon]]''}}
         |2={{clade
            |1=''[[Notosuchus]]''
            |2={{clade
               |1=''[[Comahuesuchus]]''
               |2={{clade
                  |1={{clade
                     |1=''[[Chimaerasuchus]]''
                     |2=''[[Sphagesaurus]]''}}
                  |label2=Sebecosuchia
                  |2={{clade
                     |1=''[[Baurusuchus]]''
                     |2={{clade
                        |1=''[[Bretesuchus]]''
                        |2=''[[Iberosuchus]]''
}} }} }} }} }} }} }} }} }}
}}
|data3=
{{Cladogram|title='''Larsson and Sues, 2007'''<ref name=LAS07/>
|clades={{clade| style=font-size:85%;line-height:80%
|label1=Metasuchia
|1={{clade
   |label1='''Notosuchia'''
   |1={{clade
      |1=''[[Notosuchus]]''
      |2=''[[Malawisuchus]]''}}
   |2={{clade
      |1=''[[Araripesuchus]]''
      |2={{clade
         |1={{clade
            |1=''[[Baurusuchus|{{bg|#E3F5FF|''Baurusuchus''}}]]''
            |2={{clade
               |1=[[Neosuchia]]
               |label2=[[Sebecia]]
               |2={{clade
                  |1=''[[Pabwehshi|{{bg|#E3F5FF|''Pabwehshi''}}]]''
                  |2={{clade
                     |label1=[[Sebecidae|{{bg|#E3F5FF|Sebecidae}}]]
                     |1={{clade
                        |1=''[[Sebecus|{{bg|#E3F5FF|''Sebecus''}}]]''
                        |2=''[[Bretesuchus|{{bg|#E3F5FF|''Bretesuchus''}}]]''}}
                     |2=[[Peirosauridae]]
}} }} }} }} }} }} }} {{legend2|#E3F5FF}} Taxa previously assigned to Notosuchia}}
}}
}}
The clade Notosuchia has undergone many recent phylogenetic revisions. In 2000, Notosuchia was proposed to be one of two groups within the clade [[Ziphosuchia]], the other being [[Sebecosuchia]], which included deep snouted forms such as [[baurusuchid]]s and [[sebecid]]s.<ref name=OBC00>{{cite journal |last=Ortega |first=F. Z. |author2=Buscalioni, A. D. |author3= Calvo, J. O.  |year=2000 |title=A new species of ''Araripesuchus'' (Crocodylomorpha, Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina) |journal=Journal of Vertebrate Paleontology |volume=20 |issue=1 |pages=57–76 |doi=10.1671/0272-4634(2000)020[0057:ANSOAC]2.0.CO;2}}</ref> The definition of Notosuchia by Sereno et al. (2001) is similar to that of Ziphosuchia as it includes within it Sebecosuchia. Pol (2003) also includes Sebecosuchia within Notosuchia.<ref name=CB97>{{cite journal |last=Pol |first=D. |year=2003 |title=New Remains of ''Sphagesaurus huenei'' (Crocodylomorpha: Mesoeucrocodylia) from the Late Cretaceous of Brazil |journal=Journal of Vertebrate Paleontology |volume=23 |issue=4 |pages=817–831 |doi=10.1671/A1015-7|bibcode=2003JVPal..23..817P |s2cid=86592576 }}</ref> More recently, a phylogenetic analysis by Larsson and Sues (2007) resulted in the naming of a new clade, [[Sebecia]], to include sebecids and [[Peirosauridae|peirosaurids]].<ref name=LAS07>{{cite journal |last=Larsson |first=H. C. E. |author2=Sues, H.-D.  |year=2007 |title=Cranial osteology and phylogenetic relationships of ''Hamadasuchus rebouli'' (Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco |journal=Zoological Journal of the Linnean Society |volume=149 |pages=533–567 |doi=10.1111/j.1096-3642.2007.00271.x |issue=4|doi-access=free |url=https://zenodo.org/records/5429220/files/source.pdf }}</ref> Baurusuchidae was considered to be polyphyletic in this study, with ''[[Pabwehshi]]'' being a basal member of Sebecia and ''[[Baurusuchus]]'' being the sister taxon to the clade containing [[Neosuchia]] and Sebecia. Thus, Sebecosuchia was no longer within Notosuchia and not considered to be a true clade, while Notosuchia was found to be a basal clade of Metasuchia.

The following [[cladogram]] simplified after the most comprehensive analysis of notosuchians as of 2014, presented by Pol ''et al.'' in 2014. It is based mainly on the data matrix published by Pol ''et al.'' (2012) which is itself a modified version of previous analyses. Thirty-one additional characters were added from other comprehensive analyses of notosuchians, e.g. Turner and Sertich (2010), Andrade ''et al.'' (2011), Montefeltro ''et al.'' (2011), Larsson and Sues (2007), and Novas ''et al.'' (2009), and 34 characters were noval, resulting in a matrix that includes 109 [[crocodyliform]]s and outgroup taxa which are scored based on 412 morphological traits.<ref name=C.stenognathus>{{Cite journal | doi = 10.1371/journal.pone.0093105| pmid = 24695105| title = A New Notosuchian from the Late Cretaceous of Brazil and the Phylogeny of Advanced Notosuchians| journal = PLOS ONE| volume = 9| issue = 4| pages = e93105| year = 2014| last1 = Pol | first1 = D. | last2 = Nascimento | first2 = P. M. | last3 = Carvalho | first3 = A. B. | last4 = Riccomini | first4 = C. | last5 = Pires-Domingues | first5 = R. A. | last6 = Zaher | first6 = H. | pmc=3973723| bibcode = 2014PLoSO...993105P| doi-access = free}}</ref>

{{clade| style=font-size:85%;line-height:80%
|label1='''Notosuchia'''
|1={{clade
   |1={{clade
      |1={{clade
         |label1=[[Uruguaysuchidae]]
         |1={{clade
            |1=''[[Araripesuchus tsangatsangana]]''
            |2={{clade
               |1=''[[Anatosuchus]]''
               |2=''[[Araripesuchus wegeneri]]''}}
            |3={{clade
               |1=''[[Araripesuchus buitreraensis]]''
               |2={{clade
                  |1=''[[Araripesuchus gomesii]]''
                  |2={{clade
                     |1=''[[Araripesuchus patagonicus]]''
                     |2=''[[Uruguaysuchus]]''}} }} }} }}
         |2={{clade
            |1=''[[Stolokrosuchus]]''
            |2={{clade
               |label1=[[Mahajangasuchidae]]
               |1={{clade
                  |1=''[[Kaprosuchus]]''
                  |2=''[[Mahajangasuchus]]''}}
               |label2=[[Peirosauridae]]
               |2={{clade
                  |1=''[[Hamadasuchus]]''
                  |2={{clade
                     |1=''[[Gasparinisuchus]]''
                     |2=''[[Lomasuchus]]''
                     |3=''[[Montealtosuchus]]''
                     |4=''[[Uberabasuchus]]''}} }} }} }} }}
   |2={{clade
      |1=''[[Candidodon]]''
      |label2=[[Ziphosuchia]]
      |2={{clade
         |1=''[[Libycosuchus]]''
         |2={{clade
            |1=''[[Simosuchus]]''
            |2={{clade
               |1={{clade
                  |1=''[[Malawisuchus]]''
                  |2=''[[Pakasuchus]]''}}
               |2={{clade
                  |1={{clade
                     |1=''[[Morrinhosuchus]]''
                     |2={{clade
                        |1=''[[Notosuchus]]''
                        |2={{clade
                           |1=''[[Coringasuchus]]''
                           |2=''[[Labidiosuchus]]''
                           |3=''[[Mariliasuchus]]''
                           |label4=[[Sphagesauridae]]
                           |4={{clade
                              |1={{clade
                                 |1=''[[Adamantinasuchus]]''
                                 |2=''[[Yacarerani]]''}}
                              |2={{clade
                                 |1={{clade
                                    |1=''[[Caipirasuchus stenognathus]]''
                                    |2={{clade
                                       |1=''[[Caipirasuchus paulistanus]]''
                                       |2=''[[Caipirasuchus montealtensis]]''}} }}
                                 |2={{clade
                                    |1=''[[Sphagesaurus]]''
                                    |2={{clade
                                       |1=''[[Armadillosuchus]]''
                                       |2=''[[Caryonosuchus]]''}} }} }} }} }} }} }}
                  |2={{clade
                     |1=''[[Chimaerasuchus]]''
                     |2={{clade
                        |1=''[[Comahuesuchus]]''
                        |label2=[[Sebecosuchia]]
                        |2={{clade
                           |1=''[[Pabwehshi]]''
                           |label2=[[Baurusuchidae]]
                           |2={{clade
                              |1=''[[Cynodontosuchus]]''
                              |2={{clade
                                 |1=''[[Pissarrachampsa]]''
                                 |2={{clade
                                    |1=''[[Stratiotosuchus]]''
                                    |2={{clade
                                       |1=''[[Campinasuchus]]''
                                       |2={{clade
                                          |1=''[[Baurusuchus albertoi]]''
                                          |2={{clade
                                             |1=''[[Baurusuchus pachecoi]]''
                                             |2=''[[Baurusuchus salgadoensis]]''}} }} }} }} }} }}
                           |4={{clade
                              |1={{clade
                                 |1=''[[Bergisuchus]]''
                                 |2=''[[Iberosuchus]]''}}
                              |label2=[[Sebecidae]]
                              |2={{clade
                                 |1={{clade
                                    |1=''[[Lorosuchus]]''
                                    |2=''[[Pehuenchesuchus]]''}}
                                 |2={{clade
                                    |1=''[[Barinasuchus]]''
                                    |2={{clade
                                       |1={{clade
                                          |1=''[[Ayllusuchus]]''
                                          |2=''[[Bretesuchus]]'' }}
                                       |2={{clade
                                          |1=Lumbrera form
                                          |2=''[[Langstonia]]''
                                          |3=''[[Sebecus]]''
                                          |4=''[[Zulmasuchus]]'' }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }}

This cladogram represents the results of the most comprehensive analysis of notosuchian relationships to date, performed in the description of ''[[Antaeusuchus taouzensis]]'' by Nicholl ''et al.'' 2021. It is largely based on the matrix from the above Pol ''et al.'' 2014 study, but also adding character scores from Leardi ''et al.'' 2015, Fiorelli ''et al.'' 2016, Leardi ''et al.'' 2018, and Martinez ''et al.'' 2018. The final matrix consisted of 121 taxa scored for 443 morphological traits.<ref>{{cite journal | vauthors = Nicholl CS, Hunt ES, Ouarhache D, Mannion PD | title = A second peirosaurid crocodyliform from the Mid-Cretaceous Kem Kem Group of Morocco and the diversity of Gondwanan notosuchians outside South America | journal = Royal Society Open Science | volume = 8 | issue = 10 | pages = Article ID 211254 | year = 2021 |  pmid=34659786|doi = 10.1098/rsos.211254 | pmc = 8511751 | doi-access = free | bibcode = 2021RSOS....811254N }}</ref>

{{clade| style=font-size:85%;line-height:80%
|label1='''Notosuchia'''
|1={{clade
 |1={{clade
  |label1=[[Uruguaysuchidae]]
  |1={{clade
   |1=''[[Araripesuchus tsangatsangana]]''
            |2={{clade
               |1=''[[Anatosuchus]]''
               |2=''[[Araripesuchus wegeneri]]''}}
            |3={{clade
               |1=''[[Araripesuchus buitreraensis]]''
               |2={{clade
                  |1=''[[Araripesuchus gomesii]]''
                  |2={{clade
                     |1=''[[Araripesuchus patagonicus]]''
                     |2=''[[Uruguaysuchus]]''}} }} }} }}
  |2={{clade
   |label1=[[Mahajangasuchidae]]
   |1={{clade
     |1=''[[Kaprosuchus]]''
     |2=''[[Mahajangasuchus]]''}}
   |label2=[[Peirosauridae]]
    |2={{clade
    |1=''[[Stolokrosuchus]]''
    |2={{clade
     |1={{clade
      |1=''[[Bayomesasuchus]]''
      |2={{clade
       |1=''[[Hamadasuchus]]''
       |2=''[[Antaeusuchus]]''}}}}
     |2={{clade
      |1={{clade
       |1=''[[Miadanasuchus]]''
       |2=''[[Barrosasuchus]]''}}
      |2={{clade
       |1=''[[Gasparinisuchus]]''
       |2=''[[Rukwasuchus]]''
       |3={{clade
        |1=''[[Uberabasuchus]]''
        |2={{clade
         |1=''[[Lomasuchus]]''
         |2=''[[Montealtosuchus]]''}}}}}}}}}}}}}}}}
 |2={{clade
  |1=''[[Candidodon]]''
  |label2=[[Ziphosuchia]]
  |2={{clade
   |1=''[[Libycosuchus]]''
   |2={{clade
    |1=''[[Simosuchus]]''
    |2={{clade
     |1=''[[Neuquensuchus]]''
     |2={{clade
      |1=''[[Pakasuchus]]''
      |2=''[[Malawisuchus]]''}}
      |label3=[[Xenodontosuchia]]
      |3={{clade
       |label1=[[Sphagesauria]]
       |1={{clade
        |1=''[[Morrinhosuchus]]''
        |2={{clade
         |1={{clade
          |1=''[[Llanosuchus]]''
          |2=''[[Notosuchus]]''}}
         |2={{clade
          |1=''[[Labidiosuchus]]''
          |2=''[[Mariliasuchus]]''
          |label3=[[Sphagesauridae]]
          |3={{clade
           |1={{clade
            |1=''[[Adamantinasuchus]]''
            |2=''[[Yacarerani]]''}}
           |2={{clade
            |1=''[[Caipirasuchus]]''
            |2={{clade
             |1=''[[Sphagesaurus]]''
             |2={{clade
              |1=''[[Caryonosuchus]]''
              |2=''[[Armadillosuchus]]''}}}}}}}}}}}}}}
       |2={{clade
        |1=''[[Chimaerasuchus]]''
        |2={{clade
         |1=''[[Comahuesuchus]]''
         |2=''[[Razanandrongobe]]''
         |label3=[[Sebecosuchia]]
         |3={{clade
          |label1=[[Baurusuchidae]]
          |1={{clade
           |1=''[[Cynodontosuchus]]''
           |2={{clade
            |1=''[[Pissarrachampsa]]''
            |2=''[[Campinasuchus]]''
            |3={{clade
             |1=''[[Stratiosuchus]]''
             |2=''[[Baurusuchus]]''}}}}}}
          |2={{clade
           |1={{clade
            |1=''[[Bergisuchus]]''
            |2=''[[Iberosuchus]]''}}
           |label2=[[Sebecidae]]
           |2={{clade
            |1={{clade
     |1=''[[Ayllusuchus]]''
    |2=''[[Bretesuchus]]''}}
            |2={{clade
      |1=''[[Barinasuchus]]''
 |2={{clade
  |1=''[[Ogresuchus]]''
|2=''[[Sebecus]]''}}}}}}}}}}}}}}}}}}}}}}}}}}}}

==Subclades==

There is uncertainty on the precise interrelationships between different notosuchian families. This is a summary on the phylogenetic definitions of higher-level notosuchian clades from Leardi ''et al.'' 2024.<ref>{{Cite journal |last1=Leardi |first1=J. M. |last2=Pol |first2=D. |last3=Montefeltro |first3=F. |last4=Marinho |first4=T. S. |last5=Ruiz |first5=J. V. |last6=Bravo |first6=G. G. |last7=Pinheiro |first7=A. E. P. |last8=Godoy |first8=P. L. |last9=Nicholl |first9=C. S. C. |last10=Lecuona |first10=A. |last11=Larsson |first11=H. C. E. |title=Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes) |year=2024 |journal=Bulletin of Phylogenetic Nomenclature |volume=1 |issue=3 |pages=44–82 |doi=10.11646/bpn.1.3.2 |doi-access=free }}</ref>

{| class="wikitable sortable" align="center" width="100%"
! Name
! Named by
! Definition
! Notes
|-
| '''Baurusuchia'''
| Walker, 1968
| The largest clade containing ''[[Baurusuchus pachecoi]]'', but not ''[[Sebecus icaeorhinus]]'', ''[[Sphagesaurus huenei]]'', ''[[Araripesuchus gomesii]]'', ''[[Montealtosuchus arrudacamposi]]'', and ''[[Crocodylus niloticus]]''.
| Includes the family '''Baurusuchidae''' and its closest relatives.
|-
| '''Eunotosuchia'''
| Ruiz ''et al.'', 2021
| The smallest clade containing ''[[Notosuchus terrestris]]'' and ''[[Uruguaysuchus aznarezi]]'' but not ''[[Crocodylus niloticus]]''.
| Would include all known notosuchians if '''Ururguaysuchidae''' are the most basal notosuchians.
|-
| '''Notosuchia'''
| Gasparini, 1971
| The largest clade containing ''[[Notosuchus terrestris]]'', but not ''[[Crocodylus niloticus]]''.
|-
| '''Peirosauria'''
| Leardi ''et al.'', 2024
| The smallest clade containing ''[[Mahajangasuchus insignis]]'', ''[[Peirosaurus torminni]]'', ''[[Lomasuchus palpebrosus]]'', and ''[[Itasuchus jesuinoi]]'', provided that it does not include ''[[Sebecus icaeorhinus]]''.
| Would be invalid if sebecids are more closely related to peirosaurids than mahajangasuchids are.
|-
| '''Sebecia'''
| Larsson & Sues, 2007
| The largest clade containing ''[[Sebecus icaeorhinus]]'' and ''[[Lomasuchus palpebrosus]]'', but not ''[[Baurusuchus pachecoi]]'', ''[[Notosuchus terrestris]]'', and ''[[Crocodylus niloticus]]''.
| Would be invalid if sebecids are more closely related to baurusuchids than to peirosaurids.
|-
| '''Sebecosuchia'''
| Simpson, 1937
| The smallest clade containing ''[[Sebecus icaeorhinus]]'' and ''[[Baurusuchus pachecoi]]'', but not ''[[Araripesuchus gomesii]]'', ''[[Montealtosuchus arrudacamposi]]'', or ''[[Crocodylus niloticus]]''.
| Would be invalid if sebecids are more closely related to peirosaurids than to baurusuchids.
|-
| '''Sphagesauria'''
| Kellner & Campos, 2007<ref name=KC07/>
| The largest clade containing ''[[Sphagesaurus huenei]]'', but not ''[[Baurusuchus pachecoi]]'', ''[[Araripesuchus gomesii]]'', ''[[Montealtosuchus arrudacamposi]]'', ''[[Sebecus icaeorhinus]]'', and ''[[Crocodylus niloticus]]''.
| Includes '''Sphagesauridae''' and close relatives such as ''Notosuchus''.
|-
| '''Xenodontosuchia'''
| Ruiz ''et al.'', 2021
| The smallest clade containing ''[[Sphagesaurus huenei]]'' and ''[[Baurusuchus pachecoi]]'', but not ''[[Uruguaysuchus aznarezi]]'', ''[[Araripesuchus gomesii]]'', ''[[Montealtosuchus arrudacamposi]]'', and ''[[Crocodylus niloticus]]''.
| May be invalid if sebecids are closely related to peirosaurids.
|-
| '''Ziphosuchia'''
| Ortega ''et al.'', 2000
| The smallest clade containing ''[[Notosuchus terrestris]]'', ''[[Libycosuchus brevirostris]]'', ''[[Sebecus icaeorhinus]]'', and ''[[Baurusuchus pachecoi]]'', but not ''[[Lomasuchus palpebrosus]]''.
| 
|-
|}


== References ==
{{Reflist}}

{{Pseudosuchia|L.|state=autocollapse}}
{{Notosuchia|N.|state=autocollapse}}
{{Mesoeucrocodylia}}
{{Taxonbar|from=Q134397}}

[[Category:Early Cretaceous crocodylomorphs]]
[[Category:Terrestrial crocodylomorphs]]
[[Category:Notosuchia| ]]
[[Category:Bathonian first appearances]]
[[Category:Maastrichtian extinctions]]
[[Category:Miocene extinctions]]
[[Category:Taxa named by Zulma Brandoni de Gasparini]]