View source for Anacleto Formation

{{Short description|Geologic formation in Argentina}}
{{Infobox rockunit
| name           = Anacleto Formation
| image          = Fm Anacleto Auca Mahuida 2.jpg
| caption        = Outcrop of the formation at [[Auca Mahuida]]
| type           = [[Geological formation]]
| period         = Campanian
| age            = [[Campanian|early-middle Campanian]]<br />~{{fossilrange|83|74.5}}
| prilithology   = [[Mudstone]]
| otherlithology = [[Sandstone]], [[limestone]]
| namedfor       = Aguada de Anacleto
| namedby        =
| location       = [[Patagonia]]
| region         = [[Neuquén Province|Neuquén]], [[Mendoza Province|Mendoza]] & [[Río Negro Province]]s
| country        = Argentina
| coordinates    = {{coord|37.9|S|68.5|W|display=inline,title}}
| paleocoordinates = {{coord|40.8|S|53.0|W|display=inline}}
| unitof         = [[Neuquén Group]]<br />&nbsp;[[Río Colorado Subgroup]]
| subunits       =
| underlies      = [[Malargüe Group]]<br />&nbsp;[[Allen Formation]]
| overlies       = [[Bajo de la Carpa Formation]]
| thickness      = {{convert|60|-|90|m|ft|abbr=on}}
| extent         = [[Neuquén Basin]]
| area           =
| map            = {{Location map+ | Argentina
| relief = 1
| width   = 250
| float   = center
| places  =
  {{Location map~ | Argentina
  | lat_deg = -37.9
  | lon_deg = -68.5
  | mark = Lightgreen pog.svg
  | marksize = 12
  }}
}}
| map_caption    =
}}

The '''Anacleto Formation''' is a [[geologic formation]] with [[outcrop]]s in the [[Argentina|Argentine]] [[Patagonia]]n provinces of [[Mendoza Province|Mendoza]], [[Río Negro Province|Río Negro]], and [[Neuquén Province|Neuquén]]. It is the youngest formation within the [[Neuquén Group]] and belongs to the [[Río Colorado Group|Río Colorado Subgroup]]. Formerly that subgroup was treated as a formation, and the Anacleto Formation was known as the '''Anacleto Member'''.<ref>Sánchez ''et al.'' (2006)</ref>

The type locality of this formation lies {{convert|40|km|mi}} west of the city of [[Neuquén]]. At its base, the Anacleto Formation conformably overlies the [[Bajo de la Carpa Formation]], also of the Río Colorado Subgroup, and it is in turn [[unconformity|unconformably]] overlain by the [[Allen Formation]] of the younger [[Malargüe Group]].<ref>Fossa Mancini ''et al.'' (1938), Leanza ''et al.'' (2004)</ref>

The Anacleto Formation varies between {{convert|60|and|90|m|ft}} thick, and consists mainly of [[claystone]]s and [[mudstone]]s, purple and dark red in color, deposited in [[fluvial]], [[lacustrine]] and [[floodplain]] [[depositional environment|environments]]. [[Geode]]s are often found scattered throughout this formation.<ref>Leanza ''et al.'' (2004), Sánchez ''et al.'' (2006)</ref>

== Fossil content ==
{{Paleobiota-key-compact}}

The following animals are known from bones found in the Anacleto Formation:
* several species of [[lizard]]s
* several species of [[mammal]]s

===Crocodylomorphs===
{| class="wikitable" align="center"
|-
! colspan="6" align="center" | '''[[Crocodylomorph]]s reported from the Anacleto Formation'''
|-
! Genus !! Species !! Presence 
!'''Material'''!! Notes !! Images
|-
|''[[Gasparinisuchus]]''
|''G. peirosauroides''
|Cañadón Amarrillo, south [[Malargüe]] city, [[Mendoza Province]].<ref name=":9">{{Cite journal |last1=Martinelli |first1=Agustín G. |last2=Sertich |first2=Joseph J.W. |last3=Garrido |first3=Alberto C. |last4=Praderio |first4=Ángel M. |date=2012 |title=A new peirosaurid from the Upper Cretaceous of Argentina: Implications for specimens referred to Peirosaurus torminni Price (Crocodyliformes: Peirosauridae) |url=https://www.academia.edu/1964846 |journal=Cretaceous Research |volume=37 |pages=191–200 |doi=10.1016/j.cretres.2012.03.017 |bibcode=2012CrRes..37..191M |issn=0195-6671}}</ref>
|PV-CRIDC-12 (right [[premaxilla]] and [[maxilla]], isolated teeth, and part of the postcranial skeleton).<ref name=":9" />
|A [[Peirosauridae|peirosaurid]] also known from the [[Bajo de la Carpa Formation]].
|
|-
| style="background:#f3e9f3;" |''[[Peirosaurus]]''
| style="background:#f3e9f3;" |''P. torminni''
| style="background:#f3e9f3;" |Cañadón Amarrillo, south [[Malargüe]] city, [[Mendoza Province]].<ref name=":9" />
| style="background:#f3e9f3;" |PV-CRIDC-12 (right [[premaxilla]] and [[maxilla]], isolated teeth, and part of the postcranial skeleton).<ref name=":9" />
| style="background:#f3e9f3;" |Specimen reassigned to ''[[Gasparinisuchus]]''.
| style="background:#f3e9f3;" |
|-
|}

===Dinosaurs===
====Ornithischians====
{| class="wikitable" align="center"
|-
! colspan="7" align="center" | '''[[Ornithischia]]ns reported from the Anacleto Formation'''
|-
! Genus !! Species 
!Stratigraphic position!! Presence 
!'''Material'''!! Notes !! Images
|-
|''[[Gasparinisaura]]''
|''G. cincosaltensis''
|
|Cinco Saltos, [[Río Negro Province]].<ref name=":7">{{Cite journal |last1=Salgado |first1=Leonardo |last2=Coria |first2=Rodolfo A. |last3=Heredia |first3=Susana E. |date=1997 |title=New Materials of Gasparinisaura cincosaltensis (Ornithischia, Ornithopoda) from the Upper Cretaceous of Argentina |url=https://www.jstor.org/stable/1306569 |journal=Journal of Paleontology |volume=71 |issue=5 |pages=933–940 |doi=10.1017/S0022336000035861 |jstor=1306569 |bibcode=1997JPal...71..933S |s2cid=132358890 |issn=0022-3360|url-access=subscription }}</ref>
|Multiple specimens.<ref name=":7" />
|A small [[Ornithopoda|ornithopod]].
|[[File:Gasparinisaura BW.jpg|center|150 px]]
|-
|}

====Sauropods====
[[Nest]]s of [[dinosaur egg]]s, many with preserved [[embryo]]s inside, have been discovered in large quantities at the famous [[Auca Mahuevo]] locality, and have been attributed to titanosaurs.<ref>Salgado ''et al.'' (2005), Sánchez ''et al.'' (2006)</ref>

{| class="wikitable" align="center"
|-
! colspan="7" align="center" | '''[[Sauropod]]s reported from the Anacleto Formation'''
|-
! Genus !! Species 
!Stratigraphic Position!! Presence 
!'''Material'''!! Notes !! Images
|-
|''[[Antarctosaurus]]''
|''A. wichmannianus''
|
|<ref name="vonhuene1929">{{cite journal |last1=von Huene |first1=F. |year=1929 |title=Los saurisquios y ornitisquios del Cretáceo Argentino |journal=Anales del Museo de la Plata |volume=3 |issue=3 |pages=1–196 |language=Spanish}}</ref>
|
|A [[Titanosauria|titanosaur]].
|[[File:Antarctosaurus wichmannianus  Skeletal.png|frameless]]
|-
|''[[Barrosasaurus]]''
|''B. casamiquelai''
|Lower
|[[Neuquén Province]].<ref name=":6">{{Cite journal |last1=SALGADO |first1=LEONARDO |last2=CORIA |first2=RODOLFO A. |date=2009-09-07 |title=Barrosasaurus casamiquelai gen. et sp. nov., a new titanosaur (Dinosauria, Sauropoda) from the Anacleto Formation (Late Cretaceous: early Campanian) of Sierra Barrosa (Neuquén, Argentina) |url=http://dx.doi.org/10.11646/zootaxa.2222.1.1 |journal=Zootaxa |volume=2222 |issue=1 |pages=1–16 |doi=10.11646/zootaxa.2222.1.1 |issn=1175-5334|url-access=subscription }}</ref>
|Vertebrae.<ref name=":6" />
|A [[Titanosauria|titanosaur]].
|[[File:Barrosasaurus.png|center|150 px]]
|-
|''[[Chadititan]]''<ref>{{cite journal |last1=Agnolín |first1=Federico L. |last2=Motta |first2=Matías J. |last3=Garcia Marsà |first3=Jordi |last4=Aranciaga-Rolando |first4=Mauro A. |last5=Álvarez-Herrera |first5=Gerardo |last6=Chimento |first6=Nicolás R. |last7=Rozadilla |first7=Sebastian |last8=Brissón-Egli |first8=Federico |last9=Cerroni |first9=Mauricio A. |last10=Panzeri |first10=Karen M. |last11=Bogan |first11=Sergio |last12=Casadio |first12=Silvio |last13=Sterli |first13=Juliana |last14=Miquel |first14=Sergio E. |last15=Martínez |first15=Sergio |last16=Pérez |first16=Leandro M. |last17=Pol |first17=Diego |last18=Novas |first18=Fernando E. |year=2025 |title=New fossiliferous locality from the Anacleto Formation (Late Cretaceous, Campanian) from northern Patagonia, with the description of a new titanosaur |journal=[[Revista del Museo Argentino de Ciencias Naturales]] |volume=26 |issue=2 |pages=217–259 |url=http://revista.macn.gob.ar/ojs/index.php/RevMus/article/view/885/715 |doi=10.22179/REVMACN.26.885 |issn=1853-0400|doi-access=free }}</ref>
|''C. calvoi''
| Lower
|General Roca city, Río Negro Province.
| A few fragment holotype consists of caudal vertebrae, partial humeri, femur bone, ulna, and pubis.
|A [[Titanosauria|titanosaur]].
| [[File:Chadititan calvoi.png|center|150px]]
|-
|''[[Laplatasaurus]]''
|''L. araukanicus''
|Uppermost
|Several localities in Patagonia.<ref name=":8">{{Cite journal |last=Gallina |first=Pablo Ariel |date=October 2015 |title=Reassessment of Laplatasaurus araukanicus (Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina |url=https://www.researchgate.net/publication/279251784 |journal=Ameghiniana |volume=52 |issue=5 |pages=487–501 |doi=10.5710/AMGH.08.06.2015.2911|bibcode=2015Amegh..52..487G |s2cid=131595654 }}</ref>
|Multiple specimens.<ref name=":8" />
|A [[Titanosauria|titanosaur]].
|[[File:Laplatasaurus Tibia-Fibula.png|center|150 px]]
|-
|''[[Narambuenatitan]]''
|''N. palomoi''
|Lower
|
|Remains of a subadult.<ref>{{Cite web |title=A new titanosaur sauropod dinosaur from the Upper Cretaceous of North Patagonia, Argentina - Acta Palaeontologica Polonica |url=https://www.app.pan.pl/article/item/app20100019.html |access-date=2022-12-07 |website=www.app.pan.pl |language=en}}</ref>
|A [[Titanosauria|titanosaur]].
|
|-
|''[[Neuquensaurus]]''
|''N. australis''
|Uppermost
|Cinco Saltos, [[Río Negro Province]].<ref>{{Cite journal |last1=Salgado |first1=Leonardo |last2=Apesteguía |first2=Sebastián |last3=Heredia |first3=Susana E. |date=2005 |title=A New Specimen of Neuquensaurus australis, a Late Cretaceous Saltasaurine Titanosaur from North Patagonia |url=https://www.jstor.org/stable/4524482 |journal=Journal of Vertebrate Paleontology |volume=25 |issue=3 |pages=623–634 |doi=10.1671/0272-4634(2005)025[0623:ANSONA]2.0.CO;2 |jstor=4524482 |s2cid=85606492 |issn=0272-4634|url-access=subscription }}</ref>
| Six caudal vertebrae.
|A [[Saltasaurinae|saltasaurine]].
|[[File:Neuquensaurus restoration.jpg|center|150 px]]
|-
|''[[Pitekunsaurus]]''
|''P. macayai''
|
|
|"Braincase, left frontal, one tooth, four [[cervical vertebrae]], three dorsal vertebrae, four caudal vertebrae, right ulna and scapula, proximal extreme of left [[femur]], rib fragments and uncertain remains".<ref>{{Cite journal |last1=Filippi |first1=L. |last2=Garrido |first2=Alberto |date=September 2008 |title=Pitekunsaurus macayai gen. et sp. nov., New titanosaur (Saurischia, Sauropoda) From Upper Cretaceous NeuquéN Basin, Argentina |url=https://www.researchgate.net/publication/289801911 |journal=Ameghiniana |volume=45 |issue=3 |pages=575–590}}</ref>
|A [[Titanosauria|titanosaur]].
|
|-
|style="background:#FEF6E4;" |''[[Teratopodus]]''
|style="background:#FEF6E4;" |''T. malarguensis''
|style="background:#FEF6E4;" |
|style="background:#FEF6E4;" |Southern [[Mendoza Province]].<ref name=":10">{{Cite journal |last1=Tomaselli |first1=María Belén |last2=Ortiz David |first2=Leonardo Daniel |last3=González Riga |first3=Bernardo Javier |last4=Coria |first4=Juan Pedro |last5=Mercado |first5=Claudio Ramón |last6=Guerra |first6=Mauricio |last7=Sánchez Tiviroli |first7=Germán |date=2022-01-01 |title=New titanosaurian sauropod tracks with exceptionally well-preserved claw impressions from the Upper Cretaceous of Argentina |url=https://www.sciencedirect.com/science/article/pii/S019566712100238X |journal=Cretaceous Research |language=en |volume=129 |pages=104990 |doi=10.1016/j.cretres.2021.104990 |bibcode=2022CrRes.12904990T |issn=0195-6671 |s2cid=238695181|url-access=subscription }}</ref>
|style="background:#FEF6E4;" |Footprint trackways.<ref name=":10" />
|style="background:#FEF6E4;" |A [[Titanosauria|titanosaur]] [[ichnotaxon]].
|style="background:#FEF6E4;" |
|-
|}

====Theropods====
The oldest known unequivocal [[bird]] footprints from [[South America]] were discovered in the Anacleto Formation. The small footprints were tentatively assigned to the [[ichnogenus]] ''[[Aquatilavipes]]'' and might have been produced by ''[[Patagopteryx]]'' (whose fossils were only found in the Bajo de la Carpa Formation however) or some unknown [[wader]]-like bird; they lack a hind toe. ''[[Ignotornis]]'' refers to similar footprints made by larger birds with a small hind toe; they might have been left by ''[[Neuquenornis]]'', but this is also only known from the Bajo de la Carpa Formation. Footprints of these two ichnogenera have also been found elsewhere, but it must be understood that assignment to the same ichnogenus does not imply a close relatedness of the organisms that produced these traces, only a similar [[morphology (biology)|morphology]].<ref>Coria ''et al.'' (2002), Lockley ''et al.'' (2006)</ref>

Even smaller and somewhat unusual footprints assigned to ''[[Barrosopus]]'' are only known from the Anacleto Formation. They were almost certainly made by some tiny theropod, but whether this was a bird is not quite clear: the innermost front toes of the animal leaving these tracks attached in a position higher than the others. In that, and in their dimensions, they are a very close match for the odd-footed [[enantiornithine]] bird ''[[Yungavolucris brevipedalis]]'', but this is only known from the [[Maastrichtian]] [[Lecho Formation]] which is some 10 million years younger.<ref>Chiappe (1993), Coria ''et al.'' (2002), Lockley ''et al.'' (2006)</ref>

{| class="wikitable" align="center"
|-
! colspan="7" align="center" | '''[[Theropod]]s reported from the Anacleto Formation'''
|-
! Genus !! Species 
!Stratigraphic Position!! Presence 
!'''Material'''!! Notes !! Images
|-
|[[Abelisauridae]]
|Indeterminate
|Lower
|Northwest Patagonia.<ref name=":0">{{Cite journal |last1=Gianechini |first1=Federico A. |last2=Apesteguía |first2=Sebastián |last3=Landini |first3=Walter |last4=Finotti |first4=Franco |last5=Juárez Valieri |first5=Rubén |last6=Zandonai |first6=Fabiana |date=2015-05-01 |title=New abelisaurid remains from the Anacleto Formation (Upper Cretaceous), Patagonia, Argentina |journal=Cretaceous Research |language=en |volume=54 |pages=1–16 |doi=10.1016/j.cretres.2014.11.009 |bibcode=2015CrRes..54....1G |issn=0195-6671|hdl=11336/37727 |hdl-access=free }}</ref><ref name=":1">{{Cite journal |last1=Baiano |first1=Mattia A. |last2=Coria |first2=Rodolfo A. |last3=Canale |first3=Juan I. |last4=Gianechini |first4=Federico A. |date=October 2021 |title=New abelisaurid material from the Anacleto Formation (Campanian, Upper Cretaceous) of Patagonia, Argentina, shed light on the diagnosis of the Abelisauridae (Theropoda, Ceratosauria) |url= |journal=Journal of South American Earth Sciences |volume=110 |pages=103402 |doi=10.1016/j.jsames.2021.103402 |bibcode=2021JSAES.11003402B |issn=0895-9811}}</ref>
|MPCN-PV 69, consisting of a partial [[premaxilla]], fragmentary vertebrae, proximal portion of both [[Humerus|humeri]], distal portion of the [[Pubis (bone)|pubis]], and an incomplete pedal ungual.<ref name=":0" /><ref name=":1" />
|An indeterminate [[Abelisauridae|abelisaurid]].
|
|-
|''[[Abelisaurus]]''
|''A. comahuensis''
|
|Lago Pellegrini stone quarries.<ref name=":2">{{Cite journal|last1=Bonaparte|first1=J.|last2=Novas|first2=E.E.|date=1985|title=Abelisaurus comahuensis, n.g., n.sp., Carnosauria del Crétacico Tardio de Patagonia|trans-title=Abelisaurus comahuensis, n.g., n.sp., Carnosauria from the Late Cretaceous of Patagonia|url=https://www.researchgate.net/publication/237714676|journal=Ameghiniana|volume=21|pages=259–265|via=ResearchGate}}</ref>
|Skull.<ref name=":2" />
|An [[Abelisauridae|abelisaurid]], originally thought to be from the [[Allen Formation]].
|[[File:Abelisaurus comahuensis jmallon.jpg|center|150 px]]
|-
|''[[Aerosteon]]''
|''A. riocoloradensis''
|
|"Cañadon Amarillo (S 37.5°, W 70.5°), north of [[Cerro Colorados|Cerro Colorado]], 1 km north of the [[Río Colorado (Argentina)|Río Colorado]] near the southern border of [[Mendoza Province]], Argentina."<ref name=":3">{{Cite journal |last1=Sereno |first1=Paul C. |last2=Martinez |first2=Ricardo N. |last3=Wilson |first3=Jeffrey A. |last4=Varricchio |first4=David J. |last5=Alcober |first5=Oscar A. |last6=Larsson |first6=Hans C. E. |date=2008-09-30 |title=Evidence for Avian Intrathoracic Air Sacs in a New Predatory Dinosaur from Argentina |journal=PLOS ONE |volume=3 |issue=9 |pages=e3303 |doi=10.1371/journal.pone.0003303 |issn=1932-6203 |pmc=2553519 |pmid=18825273|doi-access=free |bibcode=2008PLoSO...3.3303S }}</ref>
|Skeletal remains.<ref name=":3" />
|A [[Megaraptoridae|megaraptorid]].
|[[File:Aerosteon BW.jpg|center|150 px]]
|-
|''[[Aucasaurus]]''
|''A. garridoi''
|Upper
|[[Auca Mahuevo]].<ref name=":4">{{Cite journal |url=https://www.jstor.org/stable/4524238 |access-date=2022-12-07 |journal=Journal of Vertebrate Paleontology |jstor=4524238 |language=en|title=A New Close Relative of Carnotaurus sastrei Bonaparte 1985 (Theropoda: Abelisauridae) from the Late Cretaceous of Patagonia |last1=Coria |first1=Rodolfo A. |last2=Chiappe |first2=Luis M. |last3=Dingus |first3=Lowell |year=2002 |volume=22 |issue=2 |pages=460–465 |doi=10.1671/0272-4634(2002)022[0460:ANCROC]2.0.CO;2 |s2cid=131148538 |url-access=subscription }}</ref>
|Skeleton of an adult (MCF-PVPH-236).<ref name=":4" />
|An [[Abelisauridae|abelisaurid]].
|[[File:Aucasaurus garridoi by Paleocolour.jpg|center|150 px]]
|-
|[[Megaraptora]] indet.
|Indeterminate
|Upper
|[[Auca Mahuevo]].<ref name=":5">{{Cite web |last1=Baiano |first1=Mattia |last2=Coria |first2=Rodolfo |date=2018-05-14 |title=Revisiting Theropod Material from the Late Cretaceous Nesting Site Auca Mahuevo and the Possible Record of a Giant Megaraptoran |url=https://www.researchgate.net/publication/334067995 |website=ResearchGate |type=Abstract}}</ref>
|MCF-PVPH-416, a fragmentary pubic boot with unfused proximal symphysial contact.<ref name=":5" />
|A large [[megaraptora]]n.
|
|-
|}

===Squamates===
{| class="wikitable" align="center"
|-
! colspan="6" align="center" | '''[[Squamates]] reported from the Anacleto Formation'''
|-
! Genus !! Species !! Presence 
!'''Material'''!! Notes !! Images
|-
|''[[Dinilysia]]''
|''D.'' sp.
|Aguada Toledo, south of [[Mari Menuco Lake]].<ref name=":11">{{Cite journal |last1=Scanferla |first1=Carlos Agustín |last2=Canale |first2=Juan Ignacio |title=The Youngest Record of the Cretaceous Snake Genus Dinilysia (Squamata, Serpentes) |date=2007 |url=https://www.academia.edu/9863086 |journal=South American Journal of Herpetology |volume=2 |issue=1 |pages=76–81 |doi=10.2994/1808-9798(2007)2[76:TYROTC]2.0.CO;2 |s2cid=86744198 |issn=1808-9798}}</ref>
|24 articulated mid-posterior trunk vertebrae with the base of their respective ribs (UNC-CIP 1).<ref name=":11" />
|A large [[snake]].
|[[File:Dinilysia NT.jpg|center|150 px]]
|-
|}

===Testudines===
{| class="wikitable" align="center"
|-
! colspan="7" align="center" | '''[[Testudines]] reported from the Anacleto Formation'''
|-
! Genus !! Species 
!Stratigraphic Position!! Presence 
!'''Material'''!! Notes !! Images
|-
|''[[Prochelidella]]''
|''P. palomoi''
|
|Aguada Grande site, [[Neuquén Province]].<ref name=":12">{{Cite journal |last1=Maniel |first1=Ignacio J. |last2=de la Fuente |first2=Marcelo S. |last3=Filippi |first3=Leonardo F. |date=2022-09-28 |title=A new chelid turtle, Prochelidella palomoi sp. nov., from campanian Anacleto Formation, Neuquén Basin, North-Western Patagonia, Argentina |url=https://onlinelibrary.wiley.com/doi/10.1002/ar.25074 |journal=The Anatomical Record |volume=306 |issue=6 |language=en |pages=1365–1376 |doi=10.1002/ar.25074 |pmid=36169187 |s2cid=252566916 |issn=1932-8486|url-access=subscription }}</ref>
|A partially preserved skull, [[carapace]] remains, an almost complete [[plastron]], and a left [[Ilium (bone)|ilium]] (MAU-Pv-AG-452).<ref name=":12" />
|A [[Chelidae|chelid]] turtle.
|
|-
| ''[[Yaminuechelys]]''
|''Y.'' aff. ''maior''
|Lower
| [[Neuquén Province]].<ref name=":13">{{Cite journal |last1=de la Fuente |first1=Marcelo S. |last2=Maniel |first2=Ignacio J. |last3=Jannello |first3=Juan M. |last4=Filippi |first4=Leonardo S. |last5=Cerda |first5=Ignacio |date=2015-09-01 |title=Long-necked chelid turtles from the Campanian of northwestern Patagonia with comments on K/P survivorship of the genus Yaminuechelys |journal=Comptes Rendus Palevol |series=A tribute to France de Lapparent de Broin / Un hommage à France de Lapparent de Broin |language=en |volume=14 |issue=6 |pages=563–576 |doi=10.1016/j.crpv.2015.04.008 |issn=1631-0683|doi-access=free |bibcode=2015CRPal..14..563D |hdl=11336/57605 |hdl-access=free }}</ref>
|Two specimens (MAU-Pv-N-475 & MAU-Pv-PR-455).<ref name=":13" />
| A long-necked [[Chelidae|chelid]] turtle.
|
|-
|}

<!--
{| class="wikitable" align="center" width="100%"
|-
! colspan="5" align="center" |'''[[Fauna]] of the Anacleto Formation'''
|-
! Taxa
! Presence
! Description
! Images
|-
|
'''Genus:'''
* ''[[Abelisaurus]]''
# ''Placeholder placeholder''
|
|
| rowspan="99" |
[[Image:placeholder.jpg|thumb|center|200px|''[[Placeholder ]]'']]
|-
|
'''Genus:'''
* ''[[Antarctosaurus]]''
# ''Placeholder placeholder''
|
|
|-
|
'''Genus:'''
* ''[[Aucasaurus]]''
# ''Placeholder placeholder''
|
|
|-
|
'''Class:'''
* ''[[Aves]]''
# Footprints.
|
|
|-
|
'''Genus:'''
* ''[[Gasparinisaura]]''
# ''Placeholder placeholder''
|
|
|-
|
'''Suborder:'''
* ''[[Lacertilia]]''
# Multiple species.
|
|
|-
|
'''Genus:'''
* ''[[Mammalia]]''
# ''Placeholder placeholder''
|
|
|-
|
'''Genus:'''
* ''[[Pellegrinisaurus]]''
# ''Placeholder placeholder''
|
|
|-
|
'''Suborder:'''
* ''[[Theropoda]]''
# ''Placeholder placeholder''
|
|
|-
|
'''Unranked:'''
* ''[[Titanosauria]]''
# Eggs.
|
|
|-
|}
-->

== See also ==
{{Portal|Geology|Paleontology}}
* [[List of fossil sites]]
* [[Los Alamitos Formation]]
* [[Los Blanquitos Formation]]
* [[Adamantina Formation]]

{{clear}}

== References ==
{{Reflist|30em}}

=== Bibliography ===
* Chiappe, Luis M. (1993): Enantiornithine (Aves) Tarsometatarsi from the Cretaceous Lecho Formation of Northwestern Argentina. ''American Museum Novitates'' '''3083''': 1-27. [English with Spanish abstract] [http://digitallibrary.amnh.org/dspace/bitstream/2246/4981/1/N3083.pdf PDF fulltext]
* Coria, Rodolfo A.; Currie, Philip J.; Eberth, David & Garrido, Alberto (2002): Bird footprints from the Anacleto Formation (Late Cretaceous), Neuquén, Argentina. ''Ameghiniana'' '''39'''(4): 453-463. [English with Spanish abstract] [https://web.archive.org/web/20061124234955/http://www.biology.ualberta.ca/faculty/philip_currie/uploads/pdfs/2002/2002Bird_Footprints.PDF PDF fulltext]
* {{cite journal |last1=Fossa Mancini |first1=E. |author2-link=Egidio Feruglio |last2=Feruglio |first2=E. |last3=Yussen de Campana |first3=J.C. |year=1938 |title=Una reunión de geólogos de YPF y el problema de la terminología estratigráfica ("A YPF geologists' reunion and the problem of stratigraphy terminology") |journal=Boletín de Informaciones Petroleras |volume=15 |pages=1–67}}
* {{cite journal |last1=Leanza |first1=H.A. |last2=Apesteguia |first2=S. |author3-link=Fernando Novas |last3=Novas |first3=F.E. |last4=De la Fuente |first4=M.S. |year=2004 |title=Cretaceous terrestrial beds from the Neuquén Basin (Argentina) and their tetrapod assemblages |url=https://www.researchgate.net/publication/257047651 |journal=[[Cretaceous Research]] |volume=25 |issue=1 |pages=61–87 |doi=10.1016/j.cretres.2003.10.005 |bibcode=2004CrRes..25...61L |access-date=2019-02-16 |via=ResearchGate}}
* Lockley, Martin; Matsukawa, Masaki; Ohira, Hiroto; Li, Jianjun; Wright, Joanna; White, Diane & Chen, Peiji (2006): Bird tracks from Liaoning Province, China: New insights into avian evolution during the Jurassic-Cretaceous transition. ''Cretaceous Research'' '''27'''(1): 33-43. <small>{{doi|10.1016/j.cretres.2005.10.007}}</small> (HTML abstract). '''Erratum''': <small>{{doi|10.1016/j.cretres.2008.06.002}}</small>
* Salgado, L.; Coria, R.A. & Chiappe, Luis M. (2005): Osteology of the sauropod embryos from the Upper Cretaceous of Patagonia. ''Acta Palaeontologica Polonica'' '''50'''(1): 79–92. [http://www.app.pan.pl/archive/published/app50/app50-079.pdf PDF fulltext]
* {{cite journal |last1=Sánchez |first1=María Lidia |last2=Heredia |first2=Susana |last3=Calvo |first3=Jorge O. |year=2006 |title=Paleoambientes sedimentarios del Cretácico Superior de la Formación Plottier (Grupo Neuquén), Departamento Confluencia, Neuquén (Sedimentary paleoenvironments in the Upper Cretaceous Plottier Formation (Neuquen Group), Confluencia, Neuquén) |url=https://www.researchgate.net/publication/262615949 |journal=[[Revista de la Asociación Geológica Argentina]] |volume=61 |pages=3–18 |access-date=2019-02-16 |via=ResearchGate}}

== Further reading ==
* S. Brizuela and A. Albino. 2011. A Scincomorpha lizard from the Campanian of Patagonia. Cretaceous Research 32:781-785
* I. A. Cerda. 2008. Gastroliths in an ornithopod dinosaur. Acta Palaeontologica Polonica 53(2):351-355
* L. M. Chiappe, R. A. Coria, L. Dingus, F. Jackson, A. Chinsamy and M. Fox. 1998. Sauropod dinosaur embryos from the Late Cretaceous of Patagonia. Nature 396:258-261
* R. A. Coria and L. M. Chiappe. 2000. Un nuevo terópodo abelisaurio de la Fm. Río Colorado (Cretácico Superior) de la Provincia del Neuquén [A new abelisaur theropod from the Río Colorado Fm. (Upper Cretaceous) of Neuquén province]. Actas XVI Jornadas Argentinas de Paleontología de Vertebrado, San Luis, Argentina 13
* L. S. Filippi and A. C. Garrido. 2012. Nuevo registro del género ''Dinilysia'' (Squamata, Serpentes) para la Formación Anacleto (Campaniano inferior-medio), Rincón de los Sauces, Neuquén, Argentina. Ameghiniana 49(1):132-136
* B. J. González Riga. 2011. Speeds and stance of titanosaur sauropods: analysis of ''Titanopodus'' tracks from the Late Cretaceous of Mendoza, Argentina. Anais da Academia Brasileira de Ciências 83(1):279-290
* A. M. Praderio, A. G. Martinelli, and C. R. A. Candeiro. 2008. Mesoeucrocodilos en el Cretácico de Malargüe: primer registro de ''Peirosaurus tormini'' (Crocodyliformes, Peirosauridae) para la provincia de Mendoza (Argentina). Actas del 4to. Encuentro Internacional del International Center of Earth Sciences (E-ICES-4), Malargüe, Mendoza 1-7
* L. Salgado, S. Apesteguía, and S. Heredia. 2005. A new specimen of ''Neuquensaurus australis'', a Late Cretaceous saltasaurine titanosaur from North Patagonia. Journal of Vertebrate Paleontology 25(3):623-634
* L. Salgado, R. A. Coria, and S. E. Heredia. 1996. Nuevos materiales de ornitópodos (Ornithischia) en la Formación Rio Colorado (Cretácico Superior) de la Provincia de Rio Negro [New materials of ornithopods (Ornithischia) in the Rio Colorado Formation (Upper Cretaceous) of Rio Negro province]. Ameghiniana 33(4):471
* P. C. Sereno, R. N. Martínez, J. A. Wilson, D. J. Varricchio, O. A. Alcober and H. C. E. Larsson. 2008. Evidence for avian intrathoracic air sacs in a new predatory dinosaur from Argentina. PLoS ONE 3(9):e3303:1-20

[[Category:Anacleto Formation| ]]
[[Category:Geologic formations of Argentina]]
[[Category:Cretaceous Argentina]]
[[Category:Mudstone formations of Argentina]]
[[Category:Sandstone formations]]
[[Category:Limestone formations]]
[[Category:Fluvial deposits]]
[[Category:Lacustrine deposits]]
[[Category:Cretaceous paleontological sites of South America]]